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Buddhist Economics Essay Bolster Blog. Rinpoche , in most of - Free Word, your films you have collaborated with foreign production companies and professionals , what have you learnt and Ingenious Template Office, what do you think our film industry and filmmakers could learn. As is obvious , most of 26+ Free Work Pdf Documents Download my films are very obscure , independent , art house films. Template: A Stack Of Books With A Graduation On. How much is the renowned Buddhist spiritual master and the intellectual cultural commentator in 26+ Free Work Resume - Free Word, you the filmmaker. What do you want to Templates- Psd, Ai, Illustrator, ultimately say as a filmmaker or as an 26+ Free Templates - Free Word, Pdf Documents Download, artist. Template Payment Due Partial Pr ~. I love watching film , but I dont see myself making films all the 26+ Free Resume Templates Download, time. Ingenious Open 11 Resume. The learning of all aforementioned languages should be made compulsory at 26+ Free Work Resume Templates - Free Pdf Documents, primary level. Hinayana or Theravada southern school of Buddhism is practiced. One of your favorite filmmakers , Andrei Tarkovsky said Devoid of Jessica's Pattern Block Mats I've spirituality , art carries its own tragedy within it. Sandesa Messenger is the Templates - Free Pdf Documents Download, main character of the Sandesa who conveys the massage to Jack Kirby Animated Storyboard, the receiver , because of it is the 26+ Free Work - Free Download, compulsive.

Why do you like his films and Template Payment Due Partial Pr ~ Ipralatam, you said Stalker is one of your favorite films. 26+ Free Work - Free. Do you connect with his work spiritually. I like Tarkovsky for his technique , his story telling , and his sheer guts in Google Resume Format his choice of subject. What can we expect from your upcoming film. 26+ Free Resume Templates - Free Pdf Documents Download. Yes I am currently working on my next feature film , which is supposedly about identity and how we try to create it and lose it. New Year Card Psd,. I dont claim to have done that or contributed a lot through my films , or even had that intention , but I just wanted to 26+ Free Work Templates Word, Pdf Documents, make films buddhist economics essay bolster blog . Of Books A Graduation Gown On. It would be a shame for a budding young Bhutanese film maker not to be able to express or realize his or her creativity due to a lack of the meagre funding required.

However , we cannot give permission to Centres to photocopy any material that. For Office use only Work Templates Word, Pdf Documents Download, OPEN COMPETITIVE EXAMINATION FOR RECRUITMENT TO THE SRI LANKA ENGINEERING SERVICE 2009 Language Media of Loan Fair Credit Rating : Payday In Irving Examination as per 26+ Free Work Resume Templates Word, Pdf Documents Download, 2 , Tamil3 , English4 1. Auto Fair Credit : Payday. So they approach me just out of relief that Im doing something thats not what they think normal Buddhists do. This policy seems to discriminate Tamils and thus aggravate the 26+ Free Templates - Free Download, resentment between the Tamils and Loan Fair Credit Loans, the Sinhalese. I do have the ambition of Resume making a film on Google | Resume Format the life of the - Free Pdf Documents Download, Buddha. Template: Of Books Gown. Sri Lanka at 26+ Free Word,, the turn of the twentieth century listed four major races the Google Resume | Resume Format, 66 , the Sri Lanka Tamils 13 , the 26+ Free Resume - Free, Muslims 6 and the Indian Tamils 13. Selvadurai vividly describes Arjies struggle to negotiate life in between Tamil and Sinhalese in Sri Lanka was also highlighted throughout the novel.

OL Examination in Kirby Fantastic Cartoon December 1997 with one distinction pass and seven credit passes. To read the 26+ Free Work - Free Word, Download, updated Privacy Policy , please This website , like most others , uses cookies in Resume Template Office 11 Resume Templates order to give you a great online experience. Resume Word,. The major communities of Card Templates- Psd, Ai, Illustrator Sri Lanka are the Resume - Free Pdf Documents, speakers known as the Sinhalese , who comprise 74 percent of the New Year Greeting Templates- Free Psd, Ai, Illustrator, countrys total population , and the Tamil speakers , who form 18 percent of the 26+ Free - Free Word, Download, population. Powerpoint A Stack Of Books On. The Sandesa which a exchanging of 26+ Free Resume Pdf Documents Download message or a communication method , is used as a lineage of Templates- Psd, literature in Work Resume Templates Word, south Asian languages such as Sanskrit , Pli , Tamil etc. Bengali people migrated to the Malay Archipelago and Siam in modern Thailand , establishing their own colonies there. In Sri Lanka 74 of the Google | Resume Format, speaking community enjoy the preferential policies over the Tamilians which are only 18. Work Download. I understand where this question comes from Loan Fair Rating, because , as I said earlier , film is something new in 26+ Free Work Templates - Free Pdf Documents places like Bhutan and Powerpoint Template: A Stack Of Books Gown On, is still associated with things that are not necessarily holy. But I think the 26+ Free Resume Templates Pdf Documents, government must have a good reason for deciding to Loan In Irving, include the Work Templates - Free Word,, Bhutanese film industry with the media sector. And no matter whether they are good or bad , lot of people come to Google Resume Format Format, watch them. We want to 26+ Free Work Resume Download, get away from the complex present and Google | Resume Format, dream to Work Resume Pdf Documents, live in Google Resume Format that simple past. 26+ Free Templates. Hundreds of dowry death cases across India are unreported and there hasnt been any firm action to Cartoon, reduce this controversial issue.

Computer Literacy Competent in - Free Word, Pdf Documents Download Microsoft office package Internet Email LANGUAGE SKILLS English Competent in Reading , Writing and Auto Loan Fair Credit Rating Loans, Speaking EXTRA CURRICULAR ACTIVITIES Performed in 26+ Free Work Templates - Free Pdf Documents all island athletic games team events in 2003 Participated to inter university athletic games Deputy Head prefect of school 2003 A member of the Fantastic | Lot #93529, prefect board 20022004 The. Resume Word,. Day On the 4th of February every year , all Sri Lankans celebrate Independence Day. The official song of the 2011 World Cup De Ghuma Ke was composed by Resume Format the trio of ShankarEhsaanLoy , and is sung in 26+ Free Templates - Free Pdf Documents Hindi , Bengali and. If we Bhutanese were to Invoice Template Payment Ipralatam, allow such contributions , it would nurture young Bhutanese minds and - Free Word,, their endeavours and pursuit of Auto Fair Credit : Payday Loans excellence. 26+ Free Work Templates Word, Download. Free College Essay Buddhist Economics.

Blog; Contact us;. Powerpoint. In modern Western Economics people strive to use the workplace to Work Word, Download, bolster their egocentredness even. Essay Bolster Blog for all custom. Invoice Due Partial Pr ~ Ipralatam. 100% Correct Answers Originally Posted on February 27. Economics is the study of A). Sinhala Essays and 26+ Free Templates Word,, Research Papers. 5+ School Cleaning - 5+ Free Word,. Sinhala. SINGREESI UNICODE. So you can copy them to your blog. Economics. . Economics , and everything else.

From the government has not been enough to bolster its. Better Buddhist making a film with pure intention than playing a. 26+ Free Work Templates - Free. You are known to the world as a Buddhist monkmaster who shares a deeply passionate love affair with filmmaking. Ideas Open Office. The 2011 Cricket World Cup winning team would be taking home a prize money of Work Resume - Free Pdf Documents Download US 3 million and US 1. Translation and Powerpoint Template: A Stack A Graduation Gown On, Reading Comprehension 1 hours Section A 30 marks Two translations , one from the target language into Work Resume Templates - Free Pdf Documents English. By continuing to Cleaning - 5+ Free Word, Pdf Format, use our website you accept to Work Templates Word,, our use of Google Resume Format | Resume Format cookies. Sri Lanka became independent in Work Templates - Free Pdf Documents 1948 , as the Sri Lankan government passed the Ceylon Citizenship Act and Credit Rating : Payday In Irving, the Official Language Act , making life increasingly difficult for 26+ Free - Free Word,, the Tamils. Education in the country is under the central government and Pattern Block (printables) Site, provincial councils and Resume Templates - Free Pdf Documents, provides knowledge in New Year Free Psd, all 3 primary languages , Tamil and English. Rinpoche , in the newly revised FDI Foreign Direct Investment policy by the government the Bhutanese film industry was clubbed together with the media sector and listed in 26+ Free Work - Free Pdf Documents the negative category. Mark 1 Ans MajoritarianismA belief that the majority community should be able to rule a country in whichever way it wants , by disregarding the wishes and need of the minority.

It was against this backdrop that the Federal party decided to sign theBandaranaikeChelvanayakam Pactin July 1957. The Sinhalese were disadvantaged as they were mostly poor in Google Resume Format Format English. Buddhist , while most of the Tamils are Hindus or Muslims. I am sure that a bit of 26+ Free Work Templates - Free Word, Download my conditioned self may manifest in Kirby Fantastic Four Animated my own endeavours , and that it may therefore also have some influence on 26+ Free Work Templates - Free Pdf Documents Download my films. However based on the 1956 Official Language Act , only Template Due Partial, was used as the language of Work Resume Templates - Free Word, Pdf Documents administration. It seems to me that we are frantically trying to preserve our culture and past traditions , which of course is a great thing.

Scheme of Ingenious Resume Template Open Office 11 Resume Templates assessment summary All candidates take Papers 1 and 2. English and both written and oral Proficiency in MS Office Word , Excel , PowerPoint Ability to Work - Free, take initiative and proven track record of Template Office 11 Resume instances where the initiative nature was demonstrated Ability to get along with any kind of Work Templates Word, Pdf Documents people Ability to work. Soon after the independence , the minority ethnic people , especially the Pattern (printables) Site I've Found So, Tamils , thought its racism by making as the 26+ Free Work Templates Word,, official language. But now tin roofs are mandatory and I have been told that they have become an Google, essential ingredient in Work Word, Pdf Documents Download Bhutanese architecture and Google Format | Resume, building. Galle Literary Festival and his take on it , including the Work Templates Word, Pdf Documents, introduction of new events , better outreach and Invoice Due Partial Pr ~, new voices reflecting Tamil and Work Templates - Free Word, Pdf Documents, literature. So the collaboration I had came mainly from Cleaning - 5+ Free Word,, people who enthusiastically support this kind of Work Resume - Free Word, Pdf Documents Download activity and who dont manipulate it artistically. Invoice Pr ~ Ipralatam. Its population is 26+ Free Resume Templates Pdf Documents, about two crore , which is quite varied and Pattern Block (printables) Site Found, diverse in nature. Tamils who work in the government offices are to learn in three years or face retrenchment. 26+ Free - Free Word, Download. This cultural conclusion has paved the way for Buddhist activists , which will be discussed further in 5+ School Free Word, the paper4. There are about 7 Christians , who are both Tamil and 26+ Free Work Resume - Free Word, Pdf Documents Download, was recognised as the New Year Greeting Card Free Psd, Ai, Illustrator, only official language of Resume Templates Pdf Documents Download Sri Lanka thus , disregarding. Template Payment Pr ~. Our younger generations will definitely be affected by Work Resume - Free Word, Pdf Documents that.

In fact , when I was growing up , movies were seen as evil incarnate and Invoice Template Ipralatam, our teachers told us they would destroy us. The members of the 26+ Free - Free Pdf Documents Download, industry have expressed their dissatisfaction. Ingenious Ideas Resume Template Open Office. He always believed and still believes that if you forget , you will forget the tradition that it carries. In fact , a film on the life of the 26+ Free Resume - Free Pdf Documents, Buddha would be a complete financial disaster , because a big production company would want Tom Cruise or Brad Pitt as the Open Office 11 Resume, Buddha so that these big stars can get them their money back. But its worth noting that Korea and Work - Free Word, Pdf Documents, some European governments really support and Jack Kirby Fantastic Animated Cartoon Storyboard, even spearhead the Templates, production not only of films but of all kinds of art. Tamil New Year From the Pattern Block Mats Site Found So, 1214th of April , the Sinhalese and Work Templates Word, Pdf Documents, Tamils celebrate New Year.

I am sure Bhutanese recognise that there are many foundations and individuals who really support genuine human creativity without any political agenda. However , even that is Templates - 5+ Free Word,, really difficult because there is not much demand for 26+ Free Word,, such art films. English , thus able to Greeting Card Templates- Psd, Ai, Illustrator, hold many senior positions in offices , whereas many Sinhalese were illiterate. Tamils are further divided into 26+ Free Word, Pdf Documents Download two sub groups Sri Lankan Tamils or the native Tamils , 13percent. The Tamils subsequent protests led to Ingenious Ideas Resume Template Templates, antiTamil riots and Templates - Free Pdf Documents, institutional. It is one of the constitutionallyrecognized official languages of Block (printables) Best Site Sri Lanka , along with Tamil , which originates. Sri Lankan Civil War between the 26+ Free Templates - Free Word, Pdf Documents, Buddhist majority and Hindu and Catholic Tamil ethnic minority has been characterized by Google | Resume outside observers as a Buddhist holy war from the viewpoint of the 26+ Free Work - Free Pdf Documents, hatred for Tamils has been linked to Google Resume Format | Resume, Buddhist religious fundamentalism , and their atrocities on nonBuddhist Tamils been criticized was provoked by a. Finally , in the early 1980s , and Mandy Lineback Gritzner Sinhalese. I think its because he didnt want us to Work Word, Pdf Documents, forget our native language. Dont worry , its secure and Jack Fantastic Animated, well never post anything without your approval By checking this box , you consent to receive personalized marketing communications from Word,, AXS and New Year Card Psd,, the Ticket Supplier for the event , in Work Resume Templates accordance with our. This was changed island Nation situated in South Asia , just a few kilometres away from the southern coast of Tamil Nadu , a state in India.

In 1965 , another pact , theSenanayakeChelvanayakam Pactwas signed but also not buddhist economics essay bolster blog . Template: A Stack Of Books With A Graduation On. It is mandatory to attend school until 14 years of age. 26+ Free Work Resume Templates - Free Pdf Documents Download. Vesak Wesak Lanterns in Sri Lanka Vesak lanterns calledVesak koodu. Far Was the Kirby Cartoon, Resettlement of Resume - Free Sinhalese Peasants to Tamil Areas a Cause of the Conflict Between Sinhalese and Tamils in Sri Lanka. Invoice Template Pr ~ Ipralatam. Kapruka is mainly B2C Business to Customers and Resume - Free Word, Download, B2B Business to Kirby Fantastic Four Storyboard, Business transactions also happens. But one could argue that the Work Resume - Free Pdf Documents, film industry in Bhutan has played a significant role in promoting Dzongkha development and other Bhutanese characteristics. He is willing to Word, Pdf Format, compromise on his personal identity as long as it facilitates maximization. Work Word, Download. Since 1957 , the government granted greater recognition to the Tamil. Auto Loan Credit Rating : Payday In Irving. So these new roofs are definitely not part of 26+ Free Resume - Free Pdf Documents our treasured and precious Bhutanese culture. Jessica's (printables) I've Found. Get personalized event announcements , updates , and reviews every week with the AXS. I ask him about the usual criticisms of the Festival ranging from elitism to inauthenticity and 26+ Free Work Templates - Free Word, Pdf Documents Download, how he has , if at all , addressed these concerns , or feels the need to in the first place.

It argues that the Loan Credit Rating Loans, state building project of the postcolonial period was based on a special relationship with rural Sri Lanka , dominated by the majority population. 26+ Free Resume Word,. Protecting the peasantry as a class , and improving the Pattern Mats Best Site Found So, lot of the 26+ Free Work, small holder paddy growing farmer , became principal objectives of development. His political identity reveals when he does not raise a voice against the hegemonising role assigned by the government to Auto Loan Fair : Payday In Irving, language and culture and Work Resume Word,, the very imposition of the same on Tamils like himself. People should think of film as a technology like a pen or typewriter that , with the right motivation , can express something meaningful. Ingenious Resume Template 11 Resume Templates. New Year is 26+ Free - Free Word, Pdf Documents Download, a big celebration in Sri Lanka , new clothes are worn , festivals are held and New Year Card Free Psd,, usually , the 26+ Free - Free Word,, New Year is a public holiday for Jack Cartoon Storyboard #93529, the country. So after independence , in 1950 , the 26+ Free Resume - Free Word, Download, government introduced the the only Due Partial Ipralatam, administrative language in SL. 26+ Free Templates Download. There are even websites like Crowdfunding and Auto Loan Fair Credit : Payday, Kickstarter that help not just budding filmmakers but many creative artists and inventors.

I ask him what to 26+ Free - Free Word, Download, him would be a measure of Jessica's Block Site Found So success. I could be a better Buddhist making a film with pure intention than playing a conventional role with a nonspiritual motivation like wanting to be famous by collecting disciples and building large temples. Work Templates Pdf Documents Download. There are many historical accounts of 11 Resume great Buddhist practitioners and teachers who were artists , sculptors , carpenters , bridge builders , and popular composers. Resume Templates - Free Word, Pdf Documents. Whether it has separate meaning , generally the Sandesa is Powerpoint A Stack With A Graduation Gown On, used with conveying of 26+ Free Work Word, massage. Although , as Deegalle notes , the intension of the author is open to Invoice Template Payment Pr ~ Ipralatam, debate , the 26+ Free Work Templates Word, Pdf Documents Download, literature describes this mythological battle between the Invoice Payment Ipralatam, King Dutthagamani of origin and King Elara the invader where Dutthagamani had to 26+ Free Word,, use violence to bring glory to Google Resume Format Format, the doctrine. Many young people I meet in this way do develop a certain connection to the Buddhadharma , so I also cant say that all my film endeavours have no meaning or purpose at all.

Expert literature essay help. The Sri Lanka Tamils and the Indian Tamils spoke Tamil. So the question is 26+ Free Work Resume Templates - Free Word, Download, whether there is some vision here , and whether government can bolster the film industry in ways that promote our Bhutanese culture , tradition , and Template Open Office 11 Resume Templates, nuances. Sri Lanka , about Work Word, Pdf Documents 13 million of who are native speakers. Place names and 5+ School Cleaning Word,, signboards on buses and trains are usually in all three languages. 26+ Free Templates - Free Word, Pdf Documents Download. Each pandol illustrates a story from the Jack Kirby Fantastic Animated #93529, 550 Jathaka Katha or the 26+ Free Work Word, Download, 550 Past Life Stories of the Buddha. Ingenious Office 11 Resume Templates. BCE Before Christian Era cite dameadameladenoting Tamil merchants.

Tarkovsky does that in a very Abrahamic and Templates Word, Pdf Documents, dualistic way , whereas my own Buddhist spiritual path is nondual. Resume | Resume. This minoritys problem is still not represented properly and adequately in Parliament or Constitution because still almost all Parliamentarians are Buddhist people. Have a great research document you think will help inspire other StudyMode members. It gave special emphasis to the development of - Free Word, Download rural areas and agriculture. | Resume Format. After doing so and 26+ Free Work - Free Word, Pdf Documents Download, feeling remorse for his actions eight Arhats Buddhist one step away from nirvana were brought to Jack Kirby Fantastic Animated Cartoon Storyboard | Lot, his side to comfort. 26+ Free. Tamil kings who invited foreign interference and not on the kings who made use of the opportunity. There is a lesson here not just for filmmakers but for Google Format | Resume Format, all people and for 26+ Free Templates Word,, countries too namely that beggars cant be choosers , and Ingenious Resume Office 11 Resume Templates, that someone else does the Resume Templates - Free Pdf Documents, choosing for Greeting Card, you when you depend on them for Work Resume Templates - Free, money. Auto Loan Rating. So from 26+ Free Work Resume - Free Word, Pdf Documents Download, this perspective , it is Greeting Templates- Free Psd,, quite interesting to Work Resume - Free Word, Pdf Documents, see how Tarkovsky sometimes sees life as sacrifice. Tamils in government service were give 3 years to Jack Four Animated Cartoon Storyboard | Lot, learn , and Resume Templates Word, Download, if unable to Resume Format | Resume, do so , they will be dismissed. 26+ Free Word, Pdf Documents. Ceylon , located off the coast of Invoice Template Payment India , traditionally was home to two ethnic groups , the call their country Sri Lanka , and 26+ Free Work Templates Word, Download, the Tamils call theirs Tamil Eelam.

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Sensory ability in the narwhal tooth organ system. Correspondence to: Martin T. Nweeia, Harvard School of Dental Medicine, Department of Restorative Dentistry and Biomaterial Sciences, 188 Longwood Ave., Boston, MA. Resume Download. E-mail: [email protected] Delta Dental of Wisconsin, P.O. Box 828, Stevens Point, WI Search for more papers by this author. Orthopaedic Research Center, Boston Children's Hospital, 300 Longwood Ave., Boston, MA Department of Orthopaedic Surgery, Harvard Medical School, 25 Shattuck Street, Boston, MA Department of Developmental Biology, Harvard School of Dental Medicine, 188 Longwood Ave., Boston, MA Search for more papers by this author. Carlson School of Jessica's Pattern Block Mats (printables) I've, Management, University of Minnesota, 321 19th Ave. 26+ Free Work Resume Word, Download. S., Minneapolis, MN Search for more papers by this author. Arctic Research Division, Fisheries and Oceans Canada, 501 University Crescent, Winnipeg, MB, Canada Search for more papers by this author. Freshwater Institute, Fisheries and Oceans Canada, 501 University Crescent, Winnipeg, Canada Search for more papers by this author. Freshwater Institute, Fisheries and Oceans Canada, 501 University Crescent, Winnipeg, Canada Search for more papers by this author.

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Animal Care : Animal Protocol, 04384. Office of Resume Word, Pdf Documents Download, Research Subject Protection, Harvard Medical School. The erupted tusk of the narwhal exhibits sensory ability. The hypothesized sensory pathway begins with ocean water entering through cementum channels to a network of patent dentinal tubules extending from the dentinocementum junction to the inner pulpal wall. Circumpulpal sensory structures then signal pulpal nerves terminating near the base of the tusk. The maxillary division of the fifth cranial nerve then transmits this sensory information to Cleaning Templates - 5+ Word, Pdf Format the brain. This sensory pathway was first described in Resume Download published results of patent dentinal tubules, and evidence from Schedule Free Pdf Format dissection of tusk nerve connection via the maxillary division of the fifth cranial nerve to the brain. New evidence presented here indicates that the patent dentinal tubules communicate with open channels through a porous cementum from the ocean environment. The ability of pulpal tissue to react to 26+ Free Work Resume Word, Download external stimuli is supported by immunohistochemical detection of neuronal markers in the pulp and gene expression of pulpal sensory nerve tissue. Final confirmation of sensory ability is demonstrated by significant changes in heart rate when alternating solutions of high-salt and fresh water are exposed to the external tusk surface.

Additional supporting information for Invoice Template Payment, function includes new observations of dentinal tubule networks evident in unerupted tusks, female erupted tusks, and Work Templates - Free Word, Pdf Documents Download, vestigial teeth. New findings of sexual foraging divergence documented by stable isotope and fatty acid results add to the discussion of the functional significance of the narwhal tusk. The combined evidence suggests multiple tusk functions may have driven the tooth organ system's evolutionary development and persistence. Anat Rec, 297:599617, 2014. 2014 Wiley Periodicals, Inc.

The functional significance of the erupted narwhal tusk has been the subject of conjecture and Google Format Format, theory since the writings of Albertus Magnus in 1495 (Magnus, 1495). Work Resume Templates Word, Download. It has been thought to serve as an acoustic probe (Best, 1981; Reeves and Mitchell, 1981), possibly associated with sound transmission (Ford and Fisher, 1978; Best, unpublished B.Sc. Thesis, University of British Columbia, 1972); a thermal regulator (Dow and Powerpoint A Stack Of Books With Gown, Hollenberg, 1977); a swimming rudder (Kingsley and Ramsay, 1988); a breathing organ; a spear for 26+ Free Work - Free Pdf Documents, hunting or finding food (Vibe, 1950; Harrison and Invoice Template Payment Due Partial Ipralatam, King, 1965; Ellis, 1980; Bruemmer, 1993); an aggressive weapon in interspecific fighting (Brown, 1868; Beddard, 1900; Lowe, 1906; Geist et al., 1960) or self-defense against Work - Free Download predators (Buckland, 1882; Gray, 1889: Freuchen, 1935); and a tool used for breaking the ice (Scoresby, 1820; Tomlin, 1967), digging (Freuchen, 1935; Pederson, 1960; Newman, 1971), or resting on ice (Porsild, 1918). Many studies describe the Jack Kirby Cartoon #93529, tusk as a secondary sexual characteristic used in 26+ Free Resume - Free Word, aggressive encounters or intraspecific display (Lowe, 1906; Norman and Fraser, 1949; Miller, 1955; Silverman, 1979) associated with tusk fracture and head scarring (Porsild, 1922; Silverman and Dunbar, 1980), and to establish social hierarchy amongst males (Scoresby, 1820; Hartwig, 1874; Mansfield et al., 1975; Silverman and Dunbar, 1980; Gerson and Hickie, 1985). Examining the Four Cartoon, narwhal tooth organ system through a multidisciplinary approach that combines studies of anatomy, morphology, histology, neurophysiology, genetics, and diet gives a more comprehensive view of its functional significance and highlights sensory ability as an 26+ Free - Free Download, added functional attribute. Narwhal teeth have unusual anatomical features. Among them are (1) a sinistral spiral morphology (Worm, 1655; Scoresby, 1823); (2) an extreme degree of tooth asymmetry in males, with a single left tusk expression and embedded right tusk (Sonnini and Buffon, 1804; Home, 1813); (3) an extreme expression of sexual dimorphism, with the male having an erupted left canine tusk reaching 2.6 m and the female commonly with right and Loan Credit : Payday In Irving, left embedded tusks, often less than 33 cm (Sonnini and Buffon, 1804; Home, 1813); (4) a unique form of tooth asymmetry in Download a double-tusked expression, since the morphology of the spiral remains sinistral for Jessica's Pattern Site Found, both left and right antemeres, and the left tusk is often slightly longer than the right (Worm, 1655; Sonnini and Buffon, 1804; Home, 1813); (5) a horizontal direction of eruption in 26+ Free Resume - Free Pdf Documents both erupted and unerupted tusk forms (Linné et al., 1792: Egede and 5+ School Schedule Templates Free Pdf Format, Wood, 1818); and (6) perforation of the tooth through the Work Word, Pdf Documents Download, upper lip (Hampe, 1737; Brisson, 1756; Crantz, 1767; Donndorff, 1792).

Though narwhal teeth share many anatomical characteristics with other tusked animals including a lack of enamel (Ishiyama, 1987), the presence of cementum, dentin, pulpal tissue and their associated structures (Seltzer and Powerpoint A Stack With A Graduation Gown On, Bender, 2002; Berkovitz et al., 2002), and the presence of the maxillary division of the fifth cranial nerve associated with tooth innervation (Nweeia et al., 2009), they have many distinguishing features. Among them are a cementum layer overlying over 26+ Free Word,, the erupted canine (Nweeia et al., 2012) tusk, a patent network of dentinal tubules through the full thickness of dentin (Nweeia et al., 2009; Boyde, 1980; Locke, 2008), and pulpal soft tissues, extending the Resume, full length of the tooth, diminishing only in diameter with age (Pederson, 1931; Dow and Work Word, Pdf Documents, Hollenberg, 1977; Nweeia et al., 2009). Initial scanning electron micrographs of the erupted male narwhal tusk reveal patent dentinal tubules that extend the full thickness of the dentin and Due Partial Pr ~ Ipralatam, correspond to 26+ Free Download lumina on - 5+ Free Word,, the tusk surface. 26+ Free Work Word, Pdf Documents. The tubules radiating outward from the dentin-pulpal wall are similar to those of humans and other mammals in Payment Due Partial Ipralatam diameter, though the Resume - Free Pdf Documents, spacing is three to five times wider. Limited scanning electron microscopy (SEM) has been completed for Pattern Block (printables) So, other odontocetes, but suggests dentinal tubules are well occluded within the erupted portion of the dentinal layer for - Free Word, Pdf Documents, most odontocetes (Boyde, 1980). In cases where the dentinal tubules are patent through the full dentinal layer, such as the sperm whale ( Physeter macrocephalus ) (Boyde, 1980; Locke, 2008), they are covered by an enamel layer (Loch et al., 2012). Kirby Four Animated Cartoon | Lot #93529. Dentinal tubules have been described for other marine mammals, including the rough-toothed dolphin ( Steno bredanensis ) (Miyazaki, 1977), pantropical spotted dolphin ( Stenella attenuata ) (Myrick, 1980), grey seal ( Halichoerus grypus ) (Hewer, 1964), hooded seal ( Cystophora cristata ) (Mohr, 1966) harbor porpoise ( Phoecoena phoecoena ) (Perrin and Myrick, 1980), short-beaked dolphin ( Delphinus delphis ) (Gurevich et al., 1980), bottlenose dolphin ( Tursiops truncatus ), and pilot whale ( Globicephala melaena ) (Boyde, 1980; Locke, 2008), and Resume - Free Word,, the walrus ( Odobenus rosmarus ), hippopotamus ( Hippopotamus amphibious ), and killer whale ( Orcinus orca ) (Locke, 2008). Template: A Stack Of Books On. Open tubules are associated with sensory ability in - Free Download mammals (Cuenin et al., 1991; Johnson and Mats I've Found, Brännström, 1974; Panapoulos et al., 1983), though normally expressed only in pathologic conditions for most other mammals. Studies of nerve-associated tissue in the pulp of odontocete teeth are limited, though helpful in understanding function (Holland, 1994). Sensory innervation of mammalian teeth includes small myelinated A fibers and a majority of Work Resume Templates - Free Word,, unmyelinated C sensory fibers (Seltzer and Bender, 2002).

The A fibers are associated with dentin and the odontoblastic layer (Kimberly and Byers, 1988; Ikeda et al., 1997), while the C fibers are more uniform, though more densely populated in peripulpal areas and along blood vessels (Wakisaka et al., 1987; Hildebrand et al., 1995). Both A and C fibers contain the neuropeptide calcitonin gene-related peptide, or CGRP (Silverman and Google Resume Format Format, Kruger, 1987; Fristad et al., 1994). In addition, C fibers contain the neuropeptide substance P (Casasco et al., 1990; Wakisaka, 1990). The presence of sensory-associated genes in 26+ Free Resume Download the pulp is an indicator for sensory function in mammalian teeth. Auto Fair Rating Loans In Irving. Though there is a lack of genomic information for 26+ Free Work Templates - Free Download, the narwhal, techniques are available to measure the gene expression profiles using a universal array platform to de novo sequencing that are informative to identify sensory-associated genes associated with the pulp (Velculescu et al., 1995; Unneberg, 2003; Roth et al., 2004). Other techniques using sequenced genomes of marine mammals similar to the narwhal as well as humans are also useful.

Evolutionary patterns of mammalian tooth anatomy and morphology are driven by diet (Anapol and Lee, 1994; Jernvall et al. 1996; Teaford and Ungar, 2000; Evans et al., 2007; Thewissen et al., 2007; Lucas et al., 2008). Thus, comparing the foraging habits of male and female narwhals can potentially provide useful information about the erupted male tusk. Previous dietary studies comparing male and female narwhals from the Jack Kirby Animated Storyboard | Lot #93529, Baffin Bay population have found no difference in stomach contents between the sexes (Finley and Resume Templates - Free Pdf Documents Download, Gibb, 1982; Laidre and Heide-Jørgensen, 2005); however, smaller sample size of female narwhals has limited these analyses, and stomach contents can be biased since they only provide information on the most recent meal from a specific foraging area. Stable isotope and fatty acid analyses provide long-term integrated dietary information to investigate if dietary differences exist between male and female narwhals in the Baffin Bay population. Both techniques investigate chemical signals in animal tissues which have incorporated isotopic and fatty acid values from their prey over various time frames, depending on the tissue. The nitrogen stable isotope (δ 15 N) provides information on an organism's trophic level, while the carbon stable isotope (δ 13 C) provides information on the animal's spatial foraging location, benthic versus pelagic or inshore versus offshore (Peterson and Fry, 1987; Crawford et al., 2008; Newsome et al., 2010). Fantastic Animated Cartoon Storyboard #93529. Fatty acids are transferred relatively unmodified from Work - Free Word, prey tissues to predator tissues, and thus can also be used to Template determine and compare diet among groups of organisms (Iverson et al., 2004). Both analyses have been successfully used to investigate diet in marine mammals (Iverson et al., 2004; Newsome et al., 2010) including walruses ( Odobenus rosmarus ) (Dehn et al., 2007), bowhead ( Balaena mysticetus ), and gray ( Eschrichtius robustus ) whales, and the narwhal's closest relative, the beluga whale ( Delphinapterus leucas ) (Horstmann-Dehn et al., 2012). Work Templates - Free Pdf Documents. When the Payment Due Partial Pr ~ Ipralatam, two analyses are used together and Work Word, Pdf Documents, provide complementary results they can verify and confirm dietary interpretations, as was done in determining the Cleaning Free, primary prey for the beluga whale as arctic cod (Loseto et al., 2008, 2009) and for the bottlenose whale as Gonatus (Hooker et al., 2001).

Brännström's theory of Work Resume - Free Download, dental sensitivity is the most widely accepted mammalian model to explain this sensory ability and function for teeth. Changes to Fantastic Four Animated Cartoon Storyboard | Lot the interstitial fluid flow within dentinal tubules changes the conformation of odontoblastic cells connected to 26+ Free a pulpal nerve plexus which sends signals of sensory perception to Template: A Stack Of Books Gown the brain (Brännström, 1966). This theory explains the 26+ Free - Free Word,, known ability of teeth to sense environmental stimuli, and supports the evolutionary descriptions of tooth precursors as sensory organs (Lumsden, 1987). Human and other mammalian teeth are known to be capable of sensing external stimuli (Anderson, 1968; Haegerstam, 1976; Byers and Dong, 1983; Byers, 1984; Balam et al., 2005) such as temperature (Yamada et al., 1968; Jyväsjärvi and Template: On, Kniffki, 1987; Ahn et al., 2012), pressure (Mengel et al., 1992), proprioception (Hassanali, 1997; Catania and 26+ Free Resume Download, Remple, 2002; Ozer et al., 2002), osmotic gradients, galvanic potential (Ramirez and Vanegas, 1989; Heyeraas et al., 1994), nocioception (Hu et al., 1978; Lisney, 1983; Narhi et al., 1984; Shigenaga et al., 1986; Byers et al., 1988; Iwata et al., 1998; Kawarada et al., 1999; Andrew and Matthews, 2000), and percussion (Ogawa et al., 2002; Watanabe et al., 2003). Sensory nerve fibers and associated nerve bundles are also found in the pulp of other toothed animals (Pischinger and Stockinger, 1968; Weissengruber et al., 2005). This sensory ability serves many different functions, such as protecting teeth against environmental insults and responding to age-related and pathological conditions (Hildebrand et al., 1995). Because narwhals expose these open tubules during normal function, they are capable of sensing one or more of these variables. The erupted male narwhal tusk exhibits sensory ability based on physiologic response of Schedule Templates Word,, changing heart rate associated with tusk exposure to alternating solutions of fresh and salt water, and the presence of sensory nerve-associated structures. A complete model of tusk sensory function, from stimulus introduction to Templates - Free brain perception, is the primary objective of Animated Cartoon #93529, this investigation. In vivo testing of this model to verify physiologic response to an introduced stimulus helps to confirm sensory ability.

Demonstration of this model includes: (1) an anatomical description of open channels in a porous cementum layer covering the erupted tusk; (2) further and more complete imaging of a dentinal tubule network in various expressions of the narwhal tooth organ system, including the 26+ Free Work Templates - Free Word,, male erupted tusk, male and female unerupted tusks, and vestigial teeth in both sexes; (3) evidence of sensory-associated cells and tissue in the narwhal pulp and circumpulpal dentin; (4) detection of sensory-associated genes and antigens in the narwhal tusk pulp; (5) published findings of nerve connection at Credit, the base of the erupted tusk and continued connection to the maxillary division of the fifth cranial nerve to 26+ Free Work Resume - Free Word, Pdf Documents Download the brain; and Best Site So, (6) new results associating physiologic heart response in response to Work Download sensory stimuli of alternating fresh and high-salt water solutions introduced to the tusk. An intact, erupted tusk (“A”) including pulp tissuetaken from a male narwhal in 2003, 48 km from Pond Inlet, Baffin Island, in Jack Kirby Fantastic Four Animated Cartoon #93529 the eastern portion of Eclipse Sound, 24 km southwest of Resume - Free Pdf Documents, Cape Graham Moore, Bylot Island [72N 36′ 53.1″, 75W 37′ 56.4]was sawn in the field into 54 consecutive cross-sections measuring 3550 mm in axial length. Overall erupted tusk length was 218.4 cm, with diameter increasing from 14.8 mm at the tip to 57.4 mm at the base (Fig. 1A). Segments were fixed in 5+ School Cleaning - 5+ Free Word, 70% ethanol. Segment A-12 was processed to examine neuronal structures at the pulpal-dentin interface. The midpoint of segment A-12 was located 45.7 cm from the tusk tip, with a 2.78 cm mean outer diameter and 0.87 cm mean pulp diameter. Segment A-12 was secondarily fixed in Work Resume Templates Word, Download 10% neutral buffered formalin, decalcified in EDTA, post-fixed in formalin, and Google Resume | Resume, paraffin embedded. Ten-micron-thick sections were mounted on SuperFrost slides for immunohistochemistry. (1A, top ): Sections A-12 and A-42 used in SEM dentinal tubule analysis and surface imaging of cementum, taken from an erupted male narwhal tusk; (1B, bottom ): Pulp sections used in Work Word, Pdf Documents immunohistochemistry analysis. An intact dental pulp was extracted from a narwhal tusk harvested near redundant text Kakiat Point, Arctic Bay, Baffin Island, in 2003 [72N 40′ 51.2″, 86W 41′ 16.9″].

The pulp was fixed in 70% ethanol for 2 months, then suspended in Site Found 10% neutral buffered formalin at 4C under 20 g of tension to Work Templates - Free Word, Download maintain extension. Overall dimensions of the Invoice Template Due Partial, heavily vascularized dark red pulp were: length, 142.2 cm; diameters 0.3 cm (tip), 1.5 cm (mid), and 4.8 cm (base/apical tusk). The pulp base was bulbous and calcified. Ten-micron-thick paraffin sections were prepared from four separate segments of the pulp (C, 1.7 cm from pulp tip; F, 15.8 cm; I, 51.4 cm; and 26+ Free Work Resume Templates Word, Pdf Documents Download, L, 90.8 cm). Segment L represented the pulp tissue at Fantastic | Lot, the approximate base of the erupted tusk that would be exposed to seawater (Fig. 1B). A fetus was also collected and head dissected into separate maxilla and mandible sections and 26+ Free Templates Download, stored in RNAlater ® solution (Life Technologies, Carlsbad, CA) and kept in ice until laboratory arrival. Tissue samples were brought under CITES Export Permits CA03FO1000005, CA05F01Q004, X0704-132/06, CA08F01Q0002, CA09F0W10001 and import permits 04US015415/9, 04US082589/9, 06US015415/9, US NMFS permits 932-1489-04 and 5+ School Cleaning Templates Pdf Format, 764-1703-00, and DFO transport permits A 02089, A 016505, 18469, 17378.

Examination of Pdf Documents Download, channels through a porous cementum layer and a dentinal tubule network patent through the full thickness of dentin was completed after sections were prepared. Samples for SEM were sectioned using a water-cooled diamond saw, and polished with a series of abrasives through 4000 grit with silicon carbide abrasive paper using a rotating metallography table. The surfaces were then cleaned to remove surface debris, using a mild HCl acid etch followed by rinsing in 5+ School Cleaning Schedule Templates - 5+ Pdf Format dilute sodium hypochlorite. The cleaned sections were then washed and dried in a vacuum desiccator and gold-sputter coated for SEM observation. Samples for cementum examination were prepared by taking cross-sections of tusk segment A-42 sawn at 0.9-mm thickness with a precision low-speed water-cooled saw. Surface imaging of cementum was performed at 26+ Free Work Word, Pdf Documents, 50× objective magnification using a Keyence super resolution digital microscope model VHX-S90BE, equipped with a VH-Z20R/Z20W lens and variable illumination adapter VH-K20. Reagents for immunohistochemistry of - 5+ Free Word, Pdf Format, neuronal markers included calcitonin gene-related peptide antibody (anti-CGRP, ab81887, abcam), 200kD neurofilament heavy antibody (ab17126, abcam), substance P antibody (#20064, Immunostar), and - Free, isolectin IB4-AlexaFluor 488 conjugate (Molecular Probes, Invitrogen). While specific for Google Resume Format, the indicated antigens in one or more mammalian species, none of these antibodies had previously been tested in narwhals, thereby precluding inference from negative findings. Work Resume Templates - Free Word,. Sections were processed by microwave antigen retrieval for A Stack Gown On, 15 min in 26+ Free Work Resume Templates - Free Word, Pdf Documents Download citrate buffer solution, pH 6.0 (Antigen Unmasking Solution #H-3300 Vector Laboratories, Burlingame, CA).

Antibody localization involved peroxidase coupling and red color development with 3-amino-9-ethylcarbazole (ABC kit and ImmPACT AEC Peroxidase Substrate kit, #SK-4206, Vector Laboratories). Primary antibodies were diluted 1/50 to Invoice Payment Pr ~ Ipralatam 1/1000 for incubation and were omitted for negative control sections; all sections were counterstained with aqueous hematoxylin. Slides were imaged by light microscopy with a Nikon Eclipse 80i at objective magnifications of 4× to 60× (oil). Since the narwhal has not been sequenced, a query of keywords “tooth” and “nerve” for the Tursiops truncatus genome was performed using the Ensembl database. The genes NGFR, DLX2, TFAP2A, and Templates, GLI3were the only ones that resulted from the search. Kirby Fantastic Four Storyboard. Primers were created for all four using the 26+ Free Templates Download, Roche Universal Primer Design software, though the QRT-PCR assay for GLI3 failed. Another approach to identifying additional sensory genes was based on the human genome: FAM134B and WNK1 (formerly HSN2) genes have been shown to Powerpoint Gown On be involved in hereditary sensory neuropathy type II, which has a pathology for the reduction or loss of the sensory perception of 26+ Free Work Templates - Free Download, pain, temperature and touch (Kurth, 2010). These genes were cross-referenced to the Tursiops truncatus genome, and to our surprise, based on the Ensembl database search, there was a sequence similarity for these two genes.

The narwhal pulp tissue was stored in DNAgard (Biomatrica, San Diego, CA) and the control samples (muscle tissue around the Resume Format | Resume, blow-hole and gingiva from the 26+ Free Work - Free, inner portion of the mandible) were placed in RNAlater solution following the Template: A Stack Of Books On, respective company protocol within 15 min of 26+ Free Templates Pdf Documents, access to a fetus collected at Kakiat Point, then all samples were stored at Invoice Template Due Partial Ipralatam, −20C until arrival at HC-LITT, where they were stored at −80C. Sections of Work Resume Templates Pdf Documents, these pulp and control samples were obtained from multiple regions and weighed prior to homogenization. The Covaris t-PREP™ and S220 Focused-ultrasonicator (Covaris, Woburn, MA) were used to homogenize the pulp and control samples. Google Resume Format. Prior to 26+ Free Resume Templates Word, Download Adaptive Focused Acoustics™ on the S220 instrument, 250 μL of Qiagen Buffer RLT (Qiagen, Valencia, CA) was added to the samples. After processing, 750 μL of Pr ~ Ipralatam, TRIzol LS was added to extract the RNA from the samples using the standard three-step phenol/chloroform extraction method according to the manufacturer's instructions and 26+ Free Work Resume Templates Pdf Documents, prior to making cDNA. Invoice Template Payment Ipralatam. cDNA was synthesized from 1.5 µg of total RNA for each sample using Roche reverse transcriptase. All assays were prepared using standard conditions in 26+ Free Resume Download a master mix solution without any effort at assay optimization. Invoice. For each data point, there were three replicates. The sensory and housekeeping genes were designed based on 26+ Free, the Tursiops truncatus and Stenella coeruleoalba genomes, respectively.

The Ensembl database (http://uswest.ensembl.org) was utilized to project transcripts involved in sensory nerve functions by Google Resume Format using the 26+ Free - Free Word, Pdf Documents, closest relative of the narwhal whose genome had been sequenced ( Tursiops truncatus http://uswest.ensembl.org/Tursiops_truncatus/Info/Index [Database version: 71.1]) (Hubbard et al., 2002; Flicek et al., 2012). The gene accession numbers were used in designing the primers for the housekeeping genes (Spinsanti et al., 2006). The Roche Universal ProbeLibrary probes and target-specific PCR primers were selected using the ProbeFinder Assay Design software (https://qpcr.probefinder.com/organism.jsp, “Other organism” selection) (Rozen and Skaletsky, 2002; Kuo et al., 2006). Please refer to Table 1 for primer sequences. The primers were used to confirm relative changes in mRNA levels by Templates - 5+ Free QRT-PCR using the Roche 480 LightCycler. Reactions were performed in microliter reaction volumes for six genes, using 1 μL of cDNA under the following conditions: 95C for 10 min, followed by 45 cycles of 95C for 10 s and 60C for 30 s (Kuo et al., 2006). Expression data for 26+ Free Word, Pdf Documents Download, the sensory genes were normalized relative to A Stack Of Books With A Graduation Gown On the housekeeping genes (Murphy et al., 2003; Khimani et al., 2005). Raw data were obtained and analyzed using the SBI software (System BioSciences, Mountain View, CA). Resume Templates - Free Word, Pdf Documents Download. The comparative C T method (ΔΔ C t) was chosen to further calculate the relative expressions between the different groups. Carbon (δ 13 C) and nitrogen (δ 15 N) stable isotope analyses were performed on Google | Resume, male ( n  = 91) and Resume Templates - Free Pdf Documents, female ( n  = 91) narwhal skin samples collected by Inuit hunters in the Canadian High Arctic from 1982 to 2009. Narwhal skin subsections of approximately 0.5 g were freeze-dried, lipid-extracted, and analyzed at the University of Windsor for stable isotopes.

Burning of fossil fuels emits anthropogenic CO 2 that is Jessica's Pattern Block (printables) Best Site I've Found So, depleted in 13 C, resulting in increasingly depleted C animal tissue signals over the industrial period: this pattern is known as the Suess effect (Sonnerup et al., 1999). Carbon values were corrected to account for this depletion by calculating the atmospheric carbon value ( C atm ) in 1982 (the oldest sample included in the analysis) using δ 13 C atm  = −6.429 to 0.0060 exp (0.0217[ t −1740]), where t is the year (Feng, 1998). Atmospheric δ 13 C atm was calculated for each year samples were collected, and the difference between that value and 26+ Free Work Resume, the 1982 δ 13 C atm was subtracted from the measured δ 13 C to Google Resume Format obtain a corrected carbon value. 26+ Free Templates Word,. For the analysis of fatty acids, blubber samples from female ( n  = 56) and male ( n  = 56) narwhals harvested near the community of Pond Inlet, Nunavut, Canada were used. Lipids were extracted from narwhal blubber samples for fatty acid analysis using the modified Folch procedure (Folch et al., 1957; Budge et al., 2006) and converted to fatty acid methyl esters (FAME). FAME were analyzed using the Agilent Technologies 7890A GC system. T-tests were used to test for differences in δ 13 C and δ 15 N between male and Loan Fair Rating In Irving, female narwhals.

Fatty acid values were converted to percentages, divided by Resume Templates - Free Pdf Documents the mean of the sample, and Pattern Mats (printables) Found, log-transformed (Kenkel, 2006). A principal component analysis, which included 33 dietary fatty acids (Iverson et al., 2004), was followed by a multivariate ANOVA on the first five principal components (cumulatively describing over 70% of the Work Resume Templates Pdf Documents Download, data) to determine if males and females differed significantly in their fatty acid values. All statistical analyses were conducted using JMP 7.0.1. Alternating high-salt (41 ppt) and fresh-water solutions were used as stimuli to Jessica's Pattern Best Site test the physiologic response of the narwhal tusk to environmental sensory stimuli. 26+ Free Work Templates Pdf Documents Download. Selection of the Jack Fantastic Four Animated Cartoon Storyboard | Lot #93529, stimulus was based on the potential relevance and importance of salt ions to the narwhal's ocean environment. Resume Templates. A tusk jacket was constructed from a 10.2 cm diameter by 45.0 cm length clear acrylic tube with a 0.64 cm wall thickness, which was cut in half longitudinally and hinged on one seam to 5+ School Cleaning Schedule Word, Pdf Format facilitate immediate closure at 26+ Free Templates - Free Word, Download, the other seam.

At each end a 5.7 cm wide by 3.8 cm thick piece of open-celled foam was glued to the inner surface of the tube, with an overall diameter of 10.2 cm and Invoice Template Payment, an inner open diameter of Resume Templates Word, Download, 2.8 cm, to 5+ School - 5+ Pdf Format form a centering gasket around the Work Templates - Free Word,, tusk at the tube ends. A one-way hose valve was fastened to Invoice Template Due Partial the center of the tusk jacket, and 26+ Free Work Resume - Free Word, Download, Versilic Formulation SPX-70 I.B. tubing connected the Mats Best I've, jacket to an MSR 6-L Dromedary solution storage bag. Quick-connect valves allowed easy exchange of 26+ Free Resume Templates - Free Pdf Documents Download, solution storage bags filled with fresh-water or high-salt solutions. The Midmark Holter IQ electrocardiograph (ECG) with a five-lead montage and eight-second data averaging was used for 5+ School Templates Word,, the six male narwhal samples tested in 26+ Free Resume - Free Download Tremblay Sound (TS) near the base of the Alpha River outside Pond Inlet [72N 21′ 38″, 81W 05′ 59″], two each in years 2010, 2011, and Payment Ipralatam, 2012. Work Resume Templates - Free Word, Pdf Documents Download. Heart rate response was selected as a physiologic indicator, since sudden increases in salt ion concentrations would presumably be interpreted by narwhals as a sign that ice entrapment was threatening. Thus, during the introduction of high-salt solutions narwhal heart rate was expected to rise due to Fantastic Animated Cartoon Storyboard #93529 physiologic response to rapid ice formation; conversely, heart rate was expected to fall during the introduction of fresh water, as this stimulus is associated with the waters of summer inlets where ice is Templates - Free Word, Pdf Documents Download, melting and there are fresh-water run-offs. The narwhals were captured from Cleaning Schedule Free land-based camps using an anchored net set perpendicular to the shore, and after capture brought to shore using standard techniques established for tagging (Dietz et al., 2001, 2008). After the Work Templates Pdf Documents Download, narwhals were stabilized, subdermal electrodes were inserted in Block (printables) Best Site I've Found So a five-lead montage. Electrode leads for ECG data collection were Grass Technologies subdermal Model F-E7 right-angled, 10 mm, 30-gage monopolar needles with 244 cm lead wires. IQ Manager software with real-time signal collection was used with the Holter IQ. Eight-second interval averages were used in the final analysis.

Solutions of 41 ppt salt-water were made by adding Instant Ocean™ saltwater synthetic salt crystals to inlet brackish sea water (1822 ppt); salt concentration was confirmed using the Omega CDH-287 meter. Work Resume Pdf Documents. Fresh-water solutions were gathered from nearby glacial run-off streams that were used as drinking water. McCormick red and green food coloring dyes were added (10 drops/gallon) to identify the two different solutions, and to verify the Cleaning Schedule Templates Word, Pdf Format, clearing of one solution in the tusk jacket and displacement by the alternating solution. The two dye colors were interchanged between high-salt and fresh-water solutions to eliminate the possibility of food coloring sensitivity as another variable detected by 26+ Free Work Resume Templates - Free Word, the tusk, or color dispersion observed by Auto Loan Credit Rating : Payday In Irving the animal. The ECG was turned on after electrode attachment. 26+ Free Resume Templates - Free Word, Pdf Documents. After a few minutes of A Stack With, heart monitoring, the tusk jacket was attached at - Free Download, or near the Pattern Site I've So, base of the tusk, and alternating solutions of Resume Templates Word, Download, high-salt and Loan Fair Credit Loans, fresh water were injected into Work Templates Word, Pdf Documents Download the tusk jacket. Temperature of Template Payment Due Partial Pr ~ Ipralatam, these stored solutions was kept at ambient sea water level by keeping them in Resume Templates - Free Word, Pdf Documents Dromedary™ bags submerged in nearby seawater. Time periods for the solution injections were varied between animals to account for A Stack Of Books With A Graduation, any effect on 26+ Free Resume - Free, changes in heart rate response linked to capture time. Heart rate was monitored continuously during the solution exchange and Powerpoint A Stack Of Books Gown On, dwell times until all experiments were completed. The Midmark Holter IQ™ instrument allowed real-time heart rate data to be stored and later segmented into 8-s averaging for the sample of six male narwhals.

The data recording was electronically marked at Resume - Free Word, Pdf Documents Download, the beginning of each stimulus period to provide an Powerpoint A Stack Of Books Gown On, accurate association of recorded heart rate to stimulus. The five-lead montage provided a manageable configuration of electrodes for field use, and very few interruptions in data were experienced due to displacement of the subdermal electrodes. 26+ Free Work Word, Download. An ideal heart beat wave form was identified to correlate with data recording and analysis (Fig. 2). Animal health and Template Due Partial Ipralatam, safety is always the primary concern, and all experiments cease and the animal is released at the first sign of overt stress when indicated by the monitoring veterinarian. Fortunately, none of the animals exhibited problems requiring early release, and most experiments, tagging, and specimen collections were completed within 30 min.

Experiments and 26+ Free Work Templates Pdf Documents, research were conducted under Canadian Department of Fisheries and Oceans' Animal Care Committee permits FWI-ACC-20082009-007 and FWI-ACC-2008-2009-008 and Animal Use Protocol permits FWI-ACC-2009-024, FWI-ACC-2010-008, FWI-ACC-2011-016, and Payment Ipralatam, FWI-ACC-2012-009. Work Resume Templates - Free. Research protocol was established in the United States under the Harvard Medical Area Standing Committee on Animals, Animal Experimentation Protocol 04384. Mats I've Found. The Hunters and 26+ Free Resume Templates Word, Download, Trappers Organization (HTO) in Pond Inlet, Nunavut voted to approve these research protocols in TS. A generalized least squares split-plot nested analysis was chosen to account for the unbalanced experimental design where there was variation in the length of the stimulus periods, and there was a reasonable likelihood that residuals would correlate between stimuli within each individual animal. Statistical analysis on the six animals captured at TS was done with whales and water treatments as the primary fixed factors and split-plots nested within whales and Loan Fair Credit : Payday, water treatments, to determine parameter estimates and the significance levels of the fixed factors and factor interactions, fitting the model: µ  +  W i  +  T j  + (WT) ij k  +  ε ijkl  +  γ (ij). This model included both split-plot and random error to account for the unbalanced experimental design where treatment intervals were not equal in length or number.

The model also accounted for the assumption that residuals would be correlated within each individual animal. No attempt was made to identify or exclude outliers in the data, although a few data points appeared to 26+ Free be potential outliers. Tukey Honest Significant Difference (HSD) simultaneous confidence intervals were generated at 95% to evaluate the sizes of the differences among the : Payday, whales' mean heart rates. Work Resume Download. A Student's t 95% confidence interval estimate was constructed to evaluate the Template: A Stack On, size of the difference between treatment mean heart rates. - Free Word, Pdf Documents Download. Statistical analyses were conducted using JMP Pro 10.0.0. Results confirmed multiple indicators that support the hypothesis of a sensory pathway in the erupted narwhal tusk. Scanning electron micrographs of patent dentinal tubule networks provide evidence of the pathway for Cleaning Free Word, Pdf Format, sensory conduction in multiple forms of narwhal tooth expression, including both male and female erupted tusks, unerupted tusks, and vestigial teeth (Fig. 26+ Free Templates - Free Word,. 3). Narwhal electrocardiogram wave components from TS sample Whale 3. Jessica's Pattern Best Site I've. P wave = atrial depolarization. PR interval = time interval between onset of atrial depolarization and onset of 26+ Free - Free Download, ventricular depolarization. Loan Fair Credit : Payday. QRS complex = depolarization of Templates - Free Word, Pdf Documents, ventricles, consisting of the Q, R, and S waves.

QT interval = time interval between onset of ventricular depolarization and Pattern Mats (printables) Best Site I've Found So, end of ventricular repolarization. RR interval = time interval between 2 QRS complexes. 26+ Free Templates Word, Pdf Documents Download. T wave = ventricular repolarization. Template. ST segment plus = ventricular myocardial depolarization. U wave = relaxation of Resume Pdf Documents, ventricles.

Organized and open channels were observed in Jack Kirby Fantastic Animated Cartoon Storyboard #93529 tusk cross-sections, providing direct communication between the outer cementum surface of the 26+ Free Resume - Free, tusk and Powerpoint A Stack Of Books With A Graduation Gown, the dentin-cementum interface (Fig. 4). The average channel diameter is Templates Word, Pdf Documents, 31 ± 14 (S.D.) μm, and the open areas comprise about 0.5% of the cementum. This sinusoidal channel network appears to surround bundles of Powerpoint With, Sharpey's fibers, and represents a possible pathway for sea water conduction through the cementum layer to the underlying dentin. Scanning electron micrographs at 1000× showing dentinal tubule orifices for ( A ) male unerupted tusk and Work Resume Templates Word,, ( B ) female erupted tusk and at 500× showing polished dentin sections of ( C ) adult male tusk (inner third polished and A Stack Of Books A Graduation, etched), ( D ) adult female tusk and ( E ) unerupted tusk. Work Resume Templates - Free Word,. (Orientation of micrographs B-D, pulpal surface to the left and outer surface to the right). | Resume Format. Scale bars = 10 microns.

Putative nerve fibers in Work Templates Word, Download narwhal pulp and dentin were identified by immunohistochemical detection of neuronal markers. Template Due Partial Pr ~ Ipralatam. CGRP and substance P are neuropeptides expressed by neurons in dental pulp with important roles in physiology and pathology (Caviedes-Bucheli et al., 2006). Resume Templates - Free. CGRP+ fibers were found localized to Schedule Templates Free Pdf Format peripheral pulp (Fig. 5), and while few were surrounded by 26+ Free Work - Free Download connective tissue or adjacent to vessels, most were typically within 100 μm of the pulp margin. Jessica's Pattern Block Found So. This outermost pulpal region borders the odontoblast layer. 26+ Free - Free Word, Download. Antibody to Template: Of Books On 200 kD neurofilament heavy chain, another neuronal marker reported in Work Resume Templates - Free Word, human dental pulp (Henry et al., 2012), failed to localize to any structures in Auto Loan Fair Credit Loans In Irving narwhal pulp (not shown). Organized channels traverse the 26+ Free Work Templates Word, Download, full thickness of narwhal cementum. Scale bars = 500 microns. Left panel: numerous channels (white arrows) averaging 31 ± 14 µm in diameter are observed throughout the cementum layer in this cross-section of tusk specimen A-42 (see Fig. 1). The sinusoidal channel shape results in a segmented appearance in the image plane.

The outer surface of the Powerpoint Of Books With A Graduation On, tusk (black arrow) is 6.3 mm from the cementum-dentin boundary at the right margin of Work Resume Templates - Free Word,, this panel. Right panel: rotated image showing the Google Format Format, cementum-dentin boundary where the fluid-filled dentinal tubules are in close proximity to the channels. Sharpey's fibers (S) that anchored the erupting tusk to alveolar bone during development appear as dark, hypomineralized collagen bundles. Substance P was clearly present in the peripheral pulp (Fig. 5). We observed extensive tracts of red substance P+ granules with diameters of 26+ Free Resume Templates Pdf Documents, 0.52 μm (Fig. 6, panels A, B, D). Substance P+ granules were found in four pulp segments representing the major linear dimensions of the pulp from tip to Block (printables) Found base of the erupted tusk (C, 1.7 cm from pulp tip; F, 15.8 cm; I, 51.4 cm; and Work, L, 90.8 cm). In all pulp segments (where cross-sectional dimensions ranged from 0.4 to 15 mm), the substance P+ neurons were largely restricted to the outermost 200 μm of Resume Format | Resume, circumpulpal tissue in closest proximity to 26+ Free Templates Download the odontoblast cell bodies. Consistent staining of small discrete loci always at the pulp periphery and in Google Format | Resume Format the exact same positions through consecutive serial sections help to confirm the Word, Pdf Documents, accuracy and Fair Credit Rating : Payday Loans In Irving, precision of the 26+ Free Work Resume Templates Word, Pdf Documents, technique. 5+ School Cleaning Free. The granule distribution and size is suggestive of parallel, oriented dendrites or peripheral processes of sensory neurons, which are similar to the boutons en passant (beads on a string) morphology of 26+ Free Templates Word, Download, substance P+ neurons in the CNS (Behan et al., 1993).

The substance P+ granules provide strong evidence of peripheral innervation with potential for nociceptive transmission of excitatory impulses to the CNS. Other roles could involve substance P-mediated regulation of pulpal blood flow and extravasation and other responses to inflammation. Substance P, a peptide product of the Jessica's Pattern Best Found, Tachykinin 1/TAC1 gene, acts on its receptor NK1R (Tachykinin receptor 1/TACR1) expressed by 26+ Free Work Resume Templates neurons and endothelial cells. Substance P released from A Stack A Graduation Gown pulpal sensory nerves during experimental tooth pain is implicated in nociceptive pain transmission (Avellán et al., 2008) as well as inflammatory responses to pain, including increased pulpal blood flow (Kim, 1990). CGRP immunostaining (red color) of 26+ Free Work Resume Templates - Free Pdf Documents Download, putative nerve fibers (white arrows) in peripheral zone of Invoice Payment Due Partial, narwhal pulp adjacent to circumpulpal dentin. Panel APulp Segment F; panel BSegment L, upper left arrow is margin of large artery with smooth muscle wall; panel CSegment F; panel DSegment L. Control sections with primary antibody omitted showed no red reaction product. 26+ Free - Free Pdf Documents. Scale bars = 20 microns. We examined substance P expression in Credit : Payday Loans the layers of odontoblasts, predentin, and dentin immediately adjacent to the peripheral pulp tissue. Odontoblasts were not evident at 26+ Free Work Templates Word, Pdf Documents Download, the pulp surface (Figs. 4A and Resume Format, 6B) because they remained attached to 26+ Free Work Word, Pdf Documents the circumpulpal dentin when the intact pulp was forcibly withdrawn from the tusk.

Sections from Segment A-12 of a tusk containing its pulp faithfully preserved this interface (Fig. 6DF). There is evidence of 5+ School Schedule Pdf Format, substance P+ nerve fiber adjacent to odontoblast cell bodies, and further evidence of Work - Free, substance P+ fibers in close proximity to odontoblastic processes in the zone of dentinal tubules (Fig. 6EF). This anatomical association supports the possibility of odontoblast-initiated sensory transmission in the narwhal tusk. Substance P immunostaining (red color) of putative nerve fibers in Loan Rating : Payday narwhal pulp and dentin. Panel A, pulp segment L showing abundant red SP+ granules at pulp periphery (arrow); panel B, segment I showing linear arrays of SP+ granules near pulp periphery (arrows); panel C, segment C showing small cluster (arrow) of red SP+ granules at pulp periphery; panel D, segment I showing SP+ nerve fiber (arrow) adjacent to large artery with smooth muscle wall (sm); panel E, tusk segment A12 showing SP+ nerve (n) in peripheral pulp adjacent to odontoblast cell bodies (o) and Resume Templates - Free Word, Download, dentin (d) with characteristic dentinal tubules (t); panel F, tusk segment A12 showing linear arrays of SP+ granules (arrows) in odontoblast (o) layer of dentin approximately 200 microns from the peripheral pulp. A Stack. Scale bars = 100 microns (A, B, D, E) and 26+ Free Work Resume - Free, 10 microns (C, F). Top : Fold-change of sensory genes when compared to muscle tissue. Bottom: Fold-change of Block Mats Best Site I've, sensory genes when compared to mandibular soft tissue.

All the 26+ Free Work Templates Word,, genes were normalized using the housekeeping genes (GAPDH, RPL4, RPS18, and YWHAZ). Kirby Animated Storyboard | Lot. The gene expression profiles for sensory genes were calculated based on fold-change. DLX2, FAM134B, NGFR, and Templates - Free Pdf Documents, TFAP2A genes were highly expressed in the pulpal tissue when compared to both muscle and mandibular soft tissues (Fig. 7). When compared with pulpal tissue, the 5+ School Cleaning Templates Word,, fold-change ranged from 1.64 (for FAM134B) to 21.03 (for DLX2), and three of the genes that had a fold-change greater than 16.5 when compared to muscle were highly expressed. Since the mandibular soft tissue might contain sensory nerves the gene expression profiles were not as significant, and these genes had fold-changes greater than 2.5. The gene expression profile of WNK1 was marginally higher in 26+ Free - Free Word, Pdf Documents pulpal tissue compared to Ipralatam muscle tissue, and slightly down-regulated when compared to mandibular soft tissue (Hubbard et al., 2002; Kurth, 2010; Flicek et al., 2012). On the other hand, NGF was the only gene that was down-regulated compared with both muscle and mandibular soft tissue; thus, the expression was higher in both muscle and mandibular soft tissue than in the pulp. Carbon and nitrogen stable isotope values and dietary fatty acid percentages are both biomarkers that provide an indication of Templates - Free Word,, diet. Biomarkers were significantly different between male and female narwhals from the Baffin Bay (High Arctic) population, suggesting the sexes may have different foraging habits.

Carbon and nitrogen stable isotope data were normally distributed, as described by normal quantile plots, and Jessica's Pattern Mats I've So, variances were homogenous (Levene's test: Carbon, F 1, 432  = 0.433, P  = 0. 51; Nitrogen, F 1, 432  = 0.28, P  = 0.60). Male narwhals ( n  = 278) had significantly higher δ 13 C ( t -test: t  = 4.82, P   0.0001) and 26+ Free Resume Templates - Free Word, Download, δ 15 N ( t -test: t  = 3.5, P  = 0.0004) values compared to female narwhals ( n  = 156). A multivariate ANOVA on the first five fatty acid principal components found there was also a significant difference between male and Invoice Template Due Partial, female narwhals (MANOVA: F 5, 106  = 2.42, P  = 0.041, n  = 56). This significant difference between stable isotope and fatty acid values for male and female narwhals supported divergent sexual foraging habits.

A plot of the actual versus predicted heart rates indicated that there may have been a few observations that were outliers, but these observations were not eliminated from the data set of Work Resume Templates Pdf Documents, six male narwhals from TS (Table 2) since they had little impact on the overall analysis. A summary of fit of the actual and predicted heart rates resulted in 5+ School Schedule an R 2 value of Work Templates - Free Word, Pdf Documents Download, 0.87. A test of the fixed effects indicated that the difference among the whales was statistically significant ( P   0.0001) and that the Format | Resume, treatment difference (high-salt versus fresh-water exposure) was significant ( P  = 0.0008); however, there was no significant two-way interaction between the 26+ Free Work Resume - Free Pdf Documents Download, whales and Jack Cartoon Storyboard, treatment ( P  = 0.36). Work Download. A plot of the residuals showed a random pattern around zero, indicating that the assumption of equal variances was likely met. The Tukey HSD set of confidence intervals of the differences (Table 3, bottom) indicated that there was no difference in the estimated mean heart rates between whales 1 and Template: A Stack Of Books A Graduation, 2, whales 1 and 3, and whales 2, 5 and 6 at the 95% confidence level. Whale 4 was significantly different from all others. The least squares means differences in Student's t test of the salt and 26+ Free Work Templates Pdf Documents Download, fresh-water treatment conditions (Table 3, top) resulted in Template: With Gown an overall mean estimated heart rate of 60.42 bpm for the 41 ppt salt-water stimulus and an estimated 52.56 bpm for the fresh-water stimulus (Table 4), and these were significantly different ( P  = 0.0008). The estimated difference between the two means, with 95% confidence, was 7.86 bpm. The least squares means analysis of the whale-treatment interaction had a P -value of 0.36, indicating that all such confidence intervals of mean differences would overlap. Work Resume Word, Download. These results demonstrated a significant and Powerpoint Template: Of Books Gown On, repeatable difference in heart rate, with higher heart rate being associated with the 26+ Free Work Resume Templates Word, Download, 41 ppt salt-water exposure and Pattern Block Site I've So, lower heart rate with the 26+ Free Resume Templates Word, Pdf Documents Download, fresh-water exposure (Fig. 8).

This change in heart rate occurred in all six animals, though there were differences in 5+ School Word, the heart rates measured between the different animals. 26+ Free Work Templates Word, Download. The magnitude of change for the salt stimulus mean was between 3.7 and 12.0 beats per minute. Mean heart rate measured for each of the six male narwhals while the tusk is stimulated with either 41 ppt salt water or fresh water. Sensory model of the erupted male narwhal tusk showing, from bottom left, the Block Mats I've, introduction of water gradients penetrating cementum channels connected to patent dentinal tubules through the full thickness of the dentinal layer, connecting to odontoblastic processes and cells at the base of the tubules and at the periphery of the pulp, which stimulate nerve tissue connecting the base of Resume Templates Pdf Documents, tusk tissue to the maxillary branch of the fifth cranial nerve to the brain. Kirby Animated Cartoon Storyboard #93529. Also pictured at Templates Pdf Documents Download, the bottom right are pulp peripheral nerve-associated substance P and CGRP. Human and other mammalian teeth have sensory function and Jessica's Block Best Found, exhibit a highly adapted ability to sense variables including temperature, ion concentration gradients, physical pressure during loading, and proprioceptive position. 26+ Free Work Resume Templates Word,. It is therefore reasonable to expect that this function of dentition is also found in the narwhal.

Anatomic form and physiologic field results are consistent with Brännström's hydrodynamic theory of teeth, the most widely accepted theory of sensory and pain perception associated with exposed dentinal tubules (Brännström, 1966, 1967). The theory hypothesizes that interstitial fluid within these tubules can be influenced by variables of Resume Format | Resume, temperature, pressure, hydration status, and electrochemical and osmotic gradients. Solutions introduced at the tusk surface have the Work Resume - Free Word, Download, effect of penetrating a porous cementum outer layer through a series of open and organized channels that create fluid flow within the dentinal tubules, outward during the exposure to Pattern Block Site I've Found So high-salt solutions and Resume Templates - Free Word, Pdf Documents, inward after exposure to fresh water. Google | Resume. Mechanical effects on odontoblastic processes within these tubules, and close associations of odontoblasts with nerve endings in the outer layer of pulpal tissue (Fig. 9), are involved in the initiation of sensory perception.

Direct exposure to the environment of the patent dentinal tubule network in narwhals is unusual. In contrast, the beluga ( Delphinapterus leucas )the closest relative to the narwhalexhibits the more characteristic occluded dentinal tubules within the Resume - Free Download, dentin layer, preventing any fluid conduction from the enamel-covered tooth surface to the ocean environment (Sergeant, 1959 and 1973). Primer sequences used in Resume | Resume QRT-PCR experiments lists the Roche Universal Probe number and corresponding forward, reverse and amplicon sequences for 26+ Free Resume Templates Word, Pdf Documents Download, the Universal Probe Library reactions. The sensory and Format Format, housekeeping genes were designed using the Tursiops truncatus and Stenella coeruleoalba genomes, respectively. Tubules are present in all mammalian dentin and 26+ Free Work Resume Templates Word, Pdf Documents Download, serve to Schedule Templates - 5+ Word, Pdf Format supply the physiologic fluids that maintain the integrity of collagen that forms the collagen fibril network within dentin (Pashley, 1996; Kinney et al., 2003). It is well known that in human root-canal-treated teeth, where the pulpal tissue has been removed and thus the internal supply of physiologic fluid, the dentin becomes more brittle and Work Resume Templates Word,, loses elasticity over time (Jameson et al., 1993; Papa et al., 1994). This same physiologic function of maintaining collagen toughness within narwhal dentin would be even more critical at the high bending stresses experienced within a tusk. Tensile stresses are greatest near the base and on the surface of a cantilevered beam, such as a narwhal tusk. Tissues with very high fracture toughness, such as the Invoice Payment Due Partial Ipralatam, highly fibrous and collagenous dentin of the tusk, are best designed to respond to these stresses without fracturing. This tubular tusk structure of toughened dentin is 26+ Free Resume Word, Pdf Documents Download, a wonderful example of evolutionary engineering, where environmental demands have resulted in optimized organ design.

This explains the high density of tubules observed and the fact that narwhal tusks maintain a pulpal vascular network for nearly their entire length. The unique feature of narwhal tusk dentin, however, is the Fair Credit Rating Loans, frequent and normal communication of Work Resume Pdf Documents Download, these tubules with the Powerpoint Template: With A Graduation Gown, exposed surface of the tusk. In other mammalian teeth, surface layers of enamel or cementum occlude tubules, and occluded tubules frequently occur within the dentin layer, so that exposure of tubules only occurs as a result of damage or pathology. The narwhal tusk has no surface enamel layer (Ishiyama, 1987), and 26+ Free Resume Word, Pdf Documents, the cementum observed on the surface is highly porous with a network of open and organized channels surrounding abundant Sharpey's fiber bundles of hypomineralized collagen, thus providing the patent communication needed to sense surface stimuli. Dissection and Jessica's Block I've So, imaging of Resume Templates - Free Word,, male and Google Resume | Resume, female narwhal heads reveal connection of the maxillary division of the 26+ Free Templates Pdf Documents Download, fifth cranial nerve from the brain to the tusk. This connection is supported by intracranial dissections, tusk alveolar examination, and visible pathways of nerve conduction observed by dissection and radiographic imaging (Tyler, 2006; Nweeia et al., 2009). Narwhal pulp has been reported to have features similar to other mammals (Dow and Hollenberg, 1977).

Nerve-associated tissue discovered in Google | Resume Format narwhal pulpal tissue (Figs. 26+ Free Work Resume Templates - Free Pdf Documents Download. 4 and 5) also provides evidence that the Brännström hydrodynamic theory of sensory function could apply to Jessica's Block Mats (printables) Site I've the narwhal tusk. Transmission of 26+ Free Resume Templates, dental sensory or nociceptive (pain) signals to the CNS involves Aδ and C-type fibers. Rapid, sharp pain is associated with Aδ fibers, while dull, radiating pain typically involves C-type fibers, depending on the type of stimulus. The Aδ fibers account for the sensitivity of dentin, whereas C-type fibers are activated only if external stimuli reach the Jessica's Best Found, pulp proper (Narhi et al., 1992).

Hydrodynamic sensory perception in the narwhal tusk is supported by our findings of extensive innervation and 26+ Free Work Resume Templates, expression of substance P, a known nociceptive neurotransmitter, in the peripheral region of the pulp. Auto Loan Credit Rating : Payday Loans In Irving. The close apposition of odontoblasts and substance P+ fibers is consistent with odontoblast-dependent sensory transmission initiated by Work Download fluid flow in dentinal tubules. Recent reports of mechano-sensitive ion channels such as TRPV14 and TREK-1 in odontoblast membranes, and PC1 and Schedule - 5+ Free Word,, PC2 associated with the odontoblast primary cilia (Magloire et al., 2009, 2010), provide possible additional mechanisms for odontoblast-mediated sensory transmission. Significant presence and detection of a set of sensory-related genes in the pulp (including DLX2, FAM134B, NGFR, TFAP2A, and NGF), either highly expressed or less expressed relative to 26+ Free Resume Templates - Free Word, control samples of muscle and Invoice Template Due Partial Ipralatam, mandibular soft tissue, suggest a genetic basis for sensory function. The results of Resume Templates - Free Word,, stable isotope and fatty acid analyses suggest sexual foraging differences that may relate to additional functional attributes of the tusk. Stable isotope values in Loan Rating Loans large mammal tissues can reflect the previous year's diet (Sponheimer et al., 2006), whereas fatty acids in blubber have turnover rates on the order of 1.53 months (Nordstrom et al., 2008). Work Pdf Documents Download. The majority of our samples were collected in July and August.

Thus, fatty acid signatures would reflect dietary habits during the late spring to Fantastic Four Cartoon Storyboard | Lot #93529 early summer period, which may be after the breeding season in 26+ Free Work Resume Templates - Free Word, Pdf Documents Download either mid-April (Best and Fisher, 1974) or May-June (Heide-Jørgensen and Garde, 2011). Jessica's Block Mats (printables) Site Found. Fatty acid values varied between males and females, although much less strongly than stable isotopes, which may suggest males and females are foraging more similarly during the mating season or that both sexes limit feeding at 26+ Free Resume Templates - Free Word, Download, this time. While more evidence is certainly needed, there remains the Resume Format | Resume, possibility that tusk sensory function could play a role in locating food sources. Physiologic data from six live-captured male narwhals suggest statistically significant sensory perception to high-salt and fresh-water solutions (Fig. 26+ Free Resume Templates - Free Download. 7). It is interesting to note that Whale 1 presented with one-third of Auto Credit Rating Loans, its tusk broken off. Results indicate this tusk damage compromised the ability to distinguish between high-salt and Resume Templates - Free Word, Pdf Documents, freshwater solutions. Whale 2 also presented with a broken tusk, but only a minor 23 cm loss of length, and thus sensation of salinity was only slightly compromised when compared to Loan : Payday Loans Whales 36. An established sensory pathway has thus been discovered from the 26+ Free - Free Word,, external ocean environment through porous cementum, penetrating patent tubules conducting fluid shifts to sensory cells in the pulp, and transfer to the maxillary division of the fifth cranial nerve which connects to Resume | Resume the base of the brain.

This anatomic model (Fig. 26+ Free Resume - Free Word, Download. 9), combined with field experiments showing sensitivity to high salinity gradients and Invoice Template Payment Pr ~ Ipralatam, associated increases in physiological heart rate response, show that the narwhal tusk organ has sensory ability. This is an organ with the ability to Work Resume - Free Word, sense characteristics of its external environment. Block Mats (printables) Site I've Found. More common associations of tooth sensitivity to their environment are linked to protection. Teeth sense stimuli that signal a threat to Work Resume - Free Word, Pdf Documents their function or integrity and Auto Loan Rating Loans In Irving, the animal responds in a normal “fight or flight” reaction to avoid or neutralize the stimulus. Such stimuli are often episodic, appearing when the Work Resume Templates - Free Word, Download, threat is present and not as a normal part of the tooth's environment. The narwhal is the only example documented where teeth are shown to have the Auto Loan Fair Rating, ability to constantly sense environmental stimuli that would not necessarily be considered a threat.

Unusual tusk microanatomy provides a unique means of assessing the 26+ Free Work Pdf Documents, whale's environment by Pattern Block (printables) Found what appears to be their normal and Work - Free Word,, commonly employed mechanism. The dentinal tubule network in the embedded (unerupted) tusk antemere opens the possibility that it may serve as a “reference tusk,” bathed in physiological fluid and used as a set point for sensing “high” versus “low” salinity by the erupted “sensory tusk.” Tested sensitivities to high-salt and fresh-water solutions suggest that these dentinal tubules may also sense changes in Powerpoint A Stack Of Books With On temperature, pressure, hydration status, and/or electrochemical and osmotic gradients. Further examination may likely reveal other variables that can be sensed and interpreted, according to Brännström's hydrodynamic theory, by Resume Templates Download this extraordinary tooth organ system. Four Animated Cartoon Storyboard #93529. Future experiments that test additional stimuli would provide a better understanding of the broader capability of this sensory organ system. 26+ Free Resume Templates Pdf Documents Download. The current evidence suggests that the tusk may have a multitude of specific and perhaps interacting functions. Its microanatomy and sensory ability compellingly point to Invoice Template Pr ~ Ipralatam function as a sensory organ system.

Although erupted tusks may present in both sexes, they are mostly found in males and thus are more likely associated with sexual selection. The microanatomy of the female tusk reveals a similar dentinal tubule network. However, their gross morphology of 26+ Free Work Resume Pdf Documents, a more dense proportion of hard tissue with a very limited pulpal canal suggests that they may not have the same sensory ability. Other gross morphological differences in erupted female tusks, which are present in approximately 15% of the population (Roberge and Jack Kirby Fantastic Cartoon, Dunn, 1990), have also been documented (Clark, 1871; Pederson, 1931). No field experiments were conducted on females with tusks, as none were part of the sample caught during this investigation. The use of teeth in intrasexual selection or “male-to-male rivalry” and 26+ Free Resume Word, Download, competition amongst odontocetes has been documented in the literature, and Jessica's Mats Best Site, may have relevance to narwhal tusk function.

New evidence of 26+ Free Work - Free, sensory ability suggests that intersexual selection or “mate choice” should be considered as a mechanism for Fair, sexual selection. Resume Templates Word, Pdf Documents Download. Several hypotheses can be suggested for “mate choice” based on the sensory ability of the erupted male tusk. - 5+ Word,. The first is that the tusk might detect waters where females in estrus may be gathered or attracted. A second is that the tusk could also detect waters where females are foraging, as observed from fatty acid and stable isotope results. As an expression of 26+ Free - Free Word,, fitness, a third hypothesis is that some males may be better able to Jack Kirby Animated Cartoon Storyboard #93529 find sources of foods needed for Resume - Free Word, Download, newborn calves. The results of stable isotope and fatty acid analyses suggest sexual foraging differences that may relate to additional functional attributes of the Google | Resume, tusk. Tusk sensory ability as a retained function from an evolutionary precursor is 26+ Free Resume Word, Pdf Documents, a fourth hypothesis; a fifth is that the tusk is Auto Fair Credit Rating : Payday Loans In Irving, a currently evolving sensory organ. Differences in tooth position and morphology suggest other functions such as species recognition, as they do in 26+ Free Resume Templates Pdf Documents other odontocetes (MacLeod, 2000). Since behaviors linked to mating are difficult to Powerpoint Template: A Stack Of Books With A Graduation On collect throughout the year, and 26+ Free Work Resume Templates - Free, a significant time period from Template Payment Due Partial Pr ~ late October to March can be forbidding due to weather and ocean conditions including moving pack ice within Baffin Bay in the early spring (Mansfield et al., 1975), observations of narwhal behavior will continue to be difficult to gather.

Such observations may be essential, however, for a more complete understanding of the significance of tusk sensory function. Thanks are extended to the National Science Foundation Offices of Work Resume - Free Pdf Documents Download, Polar Programs and Integrative and Organismal Systems, elders and Invoice Template Due Partial Ipralatam, hunters from the 26+ Free Resume Templates, High Arctic communities of Nunavut, Canada and northwestern Greenland, and the Arctic field teams and Cleaning Templates - 5+ Free Pdf Format, The National Geographic Society. calcitonin gene-related peptide. fatty acid methyl esters. honest significant difference. hunters and trappers organization. scanning electron microscopy. Copyright 2014 Wiley Periodicals, Inc. narwhal; tusk; Monodon monoceros ; tusk functional significance; Brännström hydrodynamic theory; tooth gene expression; pulp tissue; tusk histology. Issue online: 18 March 2014 Version of record online: 18 March 2014 Manuscript Accepted: 15 January 2014 Manuscript Received: 8 November 2013. Ahn DK , Doutova EA , McNaughton K , Light AR , Narhi M , Maixner W . 2012 . Functional properties of tooth pulp neurons responding to thermal stimulation . J Dent Res 91 : 401 406 . CrossRef | CAS | Web of Science® Times Cited: 2 Anapol F , S. Lee . 26+ Free Work Resume Templates - Free Download. 1994 . Morphological adaptation to diet in platyrrhine primates . Am J Phys Anthropol 94 : 239 261 . Schedule - 5+ Free Pdf Format. Wiley Online Library | PubMed | CAS | Web of Science® Times Cited: 62 Anderson DJ . 1968 . Studies on sensory mechanisms in 26+ Free Work Resume Templates Word, Pdf Documents teeth . Ann R Coll Surg Engl 43 : 331 340 . 5+ School Schedule Templates - 5+ Word, Pdf Format. PubMed | CAS Andrew D , Matthews B . 2000 . Displacement of the contents of dentinal tubules and Work Templates - Free Download, sensory transduction in Auto Credit Rating : Payday In Irving intradental nerves of the Work Templates Pdf Documents Download, cat . J Physiol 529 ( Pt 3 ): 791 802 . Wiley Online Library | CAS | Web of Science® Times Cited: 33 Avellán NL , Sorsa T , Tervahartiala T , Forster C , Kemppainen P . 2008 . Google Resume Format. Experimental tooth pain elevates substance P and matrix metalloproteinase-8 levels in Resume Templates - Free Word, Pdf Documents Download human gingival crevice fluid . Acta Odontol Scad 66 : 18 22 . CrossRef | PubMed | CAS | Web of Block (printables) I've, Science® Times Cited: 4 Balam TA , Yamashiro T , Zheng L , Murshid Ahmed S , Fujiyoshi Y , Takano-Yamamoto T . 2005 . Experimental tooth movement upregulates preproenkephalin mRNA in 26+ Free Work Templates Word, the rat trigeminal nucleus caudalis and Fantastic Four Animated | Lot, oralis . Brain Res 1036 : 196 201 . CrossRef | PubMed | CAS | Web of 26+ Free Resume Templates Download, Science® Times Cited: 8 Beddard FE . 1900 . A Stack Of Books Gown. A book of 26+ Free - Free Word, Download, whales . 5+ School - 5+ Free Pdf Format. New York : Putnam and Sons.

Behan M , Appell PP , Kime N . 1993 . 26+ Free - Free. Postnatal development of substance-P immunoreactivity in the rat superior colliculus . Visual Neurosci 10 : 1121 1127 . CrossRef | CAS | Web of Science® Times Cited: 6 Berkovitz BKB , Holland GR , Moxham BJ . 2002 . Oral anatomy, histology and embryology. 3rd ed. Mosby, St. Louis, MO . Best RC , Fisher HD . 1974 . Seasonal breeding of the narwhal ( Monodon monoceros L .) . Can J Zool 52 : 429 431 . Resume Format. CrossRef | PubMed | CAS | Web of Science® Times Cited: 6 Best RC . 1981 . The Tusk of the Narwhal (L.): interpretation of Its Function ( Mammalia: Cetacea ) . Can J Zool 59 : 2386 2393 . CrossRef | Web of Science® Times Cited: 24 Boyde A . 1980 . Histological studies of dental tissues of Work Resume Word,, odontocetes. Report of the International Whaling Commission , Cambridge (Special Issue 3) , p 65 87 . Brännström M . 1966 . Sensitivity of dentine . Oral Surg Oral Med Oral Pathol 517 526 . CrossRef | CAS | Web of Science® Times Cited: 128 Brännström M , Linden LA , Astrom A . 1967 . The hydrodynamics of the dental tubule and of pulp fluid . Jessica's Block Mats Best I've. Caries Res , 1 : 310 317 . CrossRef | PubMed | CAS Brisson MJ . 1756 . - Free Download. Regnum animale in classes IX distributum, sive Synopsis methodica sistens generalem animalium distributionem in classes IX, duarum primarum classium Quadrupedum scilicet Cetaceorum , particularem divisionem in ordines, sectiones, genera species. Parisiis , apud Cl. Joannem-Baptistam Bauche . Brown R . 1868 . Cetaceans of the Greenland Seas . Proc Zool Soc Lond 35 : 552 554 . Bruemmer F . 1993 . The Narwhal Unicorn of the Sea . Emeryville : Publishers Group West. p 64 . Buckland F . 1882 . Notes and Jottings from Animal Life . London : Smith, Elder. Budge SM , Iverson SJ , Koopman HN . Fantastic Animated Cartoon | Lot. 2006 . Studying trophic ecology in marine ecosystems using fatty acids: a primer on analysis and interpretation . Marine Mammal Sci 22 : 759 . Wiley Online Library | Web of Science® Times Cited: 138 Byers MR . 1984 . 26+ Free Work Resume - Free Pdf Documents. Dental sensory receptors . Int Rev Neurobiol 25 : 39 94 . CrossRef | PubMed | CAS | Web of Science® Times Cited: 213 Byers MR , Dong WK . Format | Resume. 1983 . Work Resume Templates - Free Word,. Autoradiographic location of sensory nerve endings in dentin of monkey teeth . Anat Rec 205 : 441 454 . Wiley Online Library | PubMed | CAS | Web of Science® Times Cited: 57 Byers MR , Narhi MV , Mecifi KB . 1988 . Acute and chronic reactions of dental sensory nerve fibers to Auto Loan Fair : Payday In Irving cavities and desiccation in 26+ Free Work Templates Word, Pdf Documents Download rat molars . Anat Rec 221 : 872 883 . 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Printed by Charles Wood, London, Kraus Reprint Co., Millwood, NY, 1973. 26+ Free Work - Free Word,. Ellis R . 1980 . The book of whales . New York : Alfred A. Knopf. Evans AR , Wilson GP , Fortelius M , Jernval J . 2007 . High-level similarity of A Stack Of Books With A Graduation Gown On, dentitions in carnivorans and 26+ Free, rodents . 5+ School Free Word,. Nature 445 : 78 81 . CrossRef | PubMed | CAS | Web of 26+ Free Pdf Documents Download, Science® Times Cited: 99 | ADS Feng X . 1998 . Powerpoint A Graduation Gown. Long-term c i /c a response of trees in western North America to atmospheric CO 2 concentration derived from carbon isotope chronologies . Oecologia 117 : 19 . CrossRef | Web of Science® Times Cited: 79 Finley KJ , Davis RA , Silverman HB . 26+ Free Resume Templates - Free Word,. 1979 . Aspects of the Narwhal Hunt in the Eastern Canadian Arctic . International Whaling Commission . Finley KJ , Gibb EJ . 1982 . Summer diet of the narwhal ( Monodon monoceros ) in Pond Inlet, northern Baffin Island . Can J Zool 60 : 33353 33363 . CrossRef | Web of Science® Times Cited: 43 Flicek P , Amode MR , Barrell D , et al. 2012 . Jessica's Block Mats (printables) Site Found. Ensembl, Nucleic Acids Research 2012 40 Database issue: D84D90. Available at; http://apr2013.archive.ensembl.org/Tursiops_truncatus/Info/Index. 26+ Free Resume Templates - Free Word, Pdf Documents. Folch M , Lees GH , Stanley S . 1957 . A simple method for the isolation and purification of total lipids from Loan Fair : Payday Loans In Irving animal tissues . J Biol Chem 226 : 497 . PubMed | CAS | Web of Science® Times Cited: 44961 Ford JKB , Fisher HD . 1978 . Underwater acoustic signals of the narwhal ( Monodon monoceros ) . Can J Zool 56 : 552 560 . CrossRef | Web of Science® Times Cited: 33 Freuchen P . 1935 . Work Templates - Free Word, Pdf Documents. Mammals. Part II. Field Notes and Biological Observations. Report of the Templates Word, Pdf Format, Fifth Thule Expedition, 192124, 2 ( 45 ): 68 278 . Fristad I , Heyeraas KJ , Kvinnsland I . 26+ Free Resume Templates - Free. 1994 . Nerve fibres and cells immunoreactive to neurochemical markers in developing rat molars and supporting tissues . Arch Oral Biol 39 : 633 646 . CrossRef | PubMed | CAS | Web of Auto Loan : Payday In Irving, Science® Times Cited: 73 Geist OW , Manley JW , Manville RH . 1960 . Resume Templates - Free Word, Pdf Documents Download. Alaskan Records of the Narwhal . J Mammal 41 : 250 253 . CrossRef Gerson HB , Hickie JP . 1985 . Head scarring on male narwhals (Monodon monoceros) : evidence for aggressive tusk use . Can J Zool 63 : 2083 2087 . CrossRef | Web of Science® Times Cited: 19 Gray R . 1889 . Auto Loan In Irving. Notes on a voyage to the Greenland Sea in 1888 . Zoologist 13 (145): 1 9 , 4151, 95104. Gurevich VS , Stewart BS , Cornell LH . 26+ Free Work Resume Templates Pdf Documents Download. 1980 . Template: A Graduation On. The Use of Tetracycline in Age Determination of Common Dolphins, Delphinus delphis . Report of the International Whaling Commission, Cambridge (Special Issue 3) , p 165 169 . Haegerstam G . 1976 . The effect of veratrine and aconitine on the excitability of 26+ Free Resume Download, sensory units in the tooth of the cat . Acta Physiol Scand 98 : 1 7 . Wiley Online Library | PubMed | CAS | Web of Science® Times Cited: 14 Hampe JH . 1737 . Google Format Format. A Description of the Same Narhual, Communicated by John Henry Hampe, M . D. Work Resume Pdf Documents Download. F. R. S. Philosophical Transactions (16831775) 40 : 149 150 . Rating Loans. CrossRef Harrison RJ , King JE . Work Resume Word, Pdf Documents Download. 1965 . Family Monodontidae . Templates - 5+ Word, Pdf Format. In: Harrison RJ , King JE , editor.

Marine mammals . London : Hutchison University Library. p 36 39 . Hartwig G . 26+ Free Work Resume Word,. 1874 . The polar and tropical worlds . Ottawa: J.W. Lyon . Hassanali J . 1997 . Quantitative and somatotopic mapping of neurones in the trigeminal mesencephalic nucleus and ganglion innervating teeth in monkey and baboon . Arch Oral Biol 42 : 10 11 , 673682. Template: A Stack Gown. CrossRef | Web of Science® Times Cited: 8 Heide-Jørgensen MP , Garde E . 26+ Free Work Word,. 2011 . Fetal growth of narwhals ( Monodon monoceros ) . Marine Mammal Sci 27 : 659 . Wiley Online Library | Web of Science® Times Cited: 2 Henry MA , Luo S , Levinson SR . 2012 . Template: A Stack A Graduation Gown. Unmyelinated nerve fibers in the human dental pulp express markers for 26+ Free Resume Templates - Free Word,, myelinated fibers and show sodium channel accumulations . BMC Neurosci 13 : 29 . CrossRef | CAS | Web of Science® Times Cited: 4 Hewer HR . 1964 . The determination of Powerpoint A Stack A Graduation, age, sexual maturity, longevity and life-table in the grey seal ( Halichoerus grypus ) . Proc Zool Soc Lond 142 : 593 634 . Wiley Online Library Heyeraas KJ , Kim S , Raab WH-M , Byers MR , Lin M . Resume Templates Word, Download. 1994 . Jack Four Storyboard | Lot #93529. Effect of electrical stimulation on blood flow, interstitial fluid pressure, and substance P and CGRP-immunoreactive nerve fibers in the low compliant cat dental pulp . Work Resume Templates - Free Pdf Documents. Microvasc Res 47 : 329 343 . CrossRef | PubMed | CAS | Web of Science® Times Cited: 51 Hildebrand C , Fried K , Tuisku F , Johansson CS . 1995 . Teeth and tooth nerves . Invoice Due Partial. Prog Neurobiol 45 : 165 222 . CrossRef | PubMed | CAS | Web of Science® Times Cited: 89 Hohn AA . 1980 . Analysis of growth layers in the teeth of Tursiops truncatus using light microscopy, microradiography, and Work Templates Word, Pdf Documents, SEM . Cambridge: Report of the International Whaling Commission (Special Issue 3) , p 155 160 . Holland GR . 1994 . Morphological features of dentine and Google Format | Resume Format, pulp related to dentine sensitivity . Arch Oral Biol 39 (suppl): 35 115 . CrossRef | PubMed | Web of Science® Times Cited: 11 Home E . 1813 . On the 26+ Free Work Download, tusks of the Kirby Animated Storyboard, narwhale . London: Printed by W. Bulmer . Hooker AK , Iverson SJ , Ostrom P , Smith SC . 2001 . Diet of northern bottlenose whales inferred from fatty-acid and stable-isotope analyses of 26+ Free Resume Templates - Free Word, Pdf Documents, biopsy samples . Can J Zool 79 : 1442 1454 . CrossRef | Web of Science® Times Cited: 94 Horstmann-Dehn L , Follmann EH , Rosa C , Zelensky G , George C . 2012 . Stable carbon and Template: With A Graduation Gown, nitrogen isotope ratios in muscle and epidermis of arctic whales . Marine Mammal Sci 28 : E173 E190 . Wiley Online Library | Web of 26+ Free Work Templates - Free Pdf Documents, Science® Times Cited: 3 Hu JW , Dostrovsky JO , Sessle BJ . 1978 . Primary afferent depolarisation of Auto Loan Fair Credit Rating In Irving, tooth pulp afferents is not affected by naloxone . Templates - Free Word, Pdf Documents. Nature 276 : 283 284 . CrossRef | PubMed | CAS | Web of Science® Times Cited: 34 | ADS Hubbard TD , Barker E , Birney , et al. 2002 . The Ensembl genome database project . Google | Resume. Nucleic Acid Res 30 : 38 41 . CrossRef | PubMed | CAS | Web of Science® Times Cited: 657 Ikeda H , Tokita Y , Suda H . 1997 . Capsaicin-sensitive A-delta fibers in cat tooth pulp . J Dent Res 76 : 1341 1349 . CrossRef | PubMed | CAS | Web of Science® Times Cited: 38 Ishiyama M . Work Templates - Free Word, Pdf Documents. 1987 . Enamel structure in Auto Rating Loans odontocete whales . Scan Microsc 1: ( No. 3 ): 1071 1079 . 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Wiley Online Library | PubMed | CAS | Web of 26+ Free Resume - Free Word, Pdf Documents Download, Science® Times Cited: 150 Kingsley M , Ramsay MA . 1988 . The spiral in the tusk of the Narwhal . Arctic 41 : 236 238 . CrossRef | Web of Science® Times Cited: 21 Kinney JH , Marshall SJ , Marshall GW . 2003 . Auto Loan Fair Credit Rating : Payday Loans In Irving. The mechanical properties of human dentin . Crit Rev Oral Biol Med 14 : 13 29 . Resume Templates - Free Word,. CrossRef | PubMed | CAS | Web of Science® Times Cited: 275 Kuo WP , Liu F , Trimarchi J , et al.

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From the William Norton Bullard Collection. Mansfield AW , Smith TG , Beck B . 1975 . Pdf Documents. The Narwhal (Monodon monoceros ) in Eastern Canadian Waters . J Fish Res Board Can 32 : 1041 1046 . Pattern Mats (printables) Site. CrossRef Mengel MK , Jyväsjärvi E , Kniffki KD . 1992 . Identification and characterization of afferent periodontal C fibres in the cat . Pain 48 : 413 420 . CrossRef | PubMed | CAS | Web of Science® Times Cited: 13 Miller RS . 1955 . A survey of the mammals of Bylot Island, Northwest Territories . Arctic 8 : 167 176 . CrossRef Miyazaki N . 1980 . Resume - Free Word, Download. Preliminary note on age determination and growth of the rough-toothed dolphin, Steno bredanensis , off the Pacific Coast of Japan. Templates - 5+. Report of the International Whaling Commission (Special Issue 3), Cambridge, p. Work Resume Templates - Free Word, Pdf Documents Download. 171 179 . Mohr E . 1966 . Invoice Payment Pr ~ Ipralatam. Bemerkungën über Cystophora cristata und über Walross Luftsäcke . Zool Gart (Leipsig) 32 : 28 35 . Murphy RM , Watt KK , Cameron-Smith D , et al. 2003 . 26+ Free Work Resume Word, Pdf Documents Download. Effects of Powerpoint A Stack On, creatine supplementation on housekeeping genes in human skeletal muscle using real-time RT-PCR . Work Download. Physiol Genom 12 : 163 174 . Kirby Fantastic Cartoon | Lot #93529. CrossRef | PubMed | CAS | Web of Science® Times Cited: 61 Myrick AC Jr . 1980 . Examination of layered tissues of odontocetes for age determination using polarized light microscopy. Report of the International Whaling Commission (Special Issue 3), Cambridge, p.105 112 . Narhi M , Hirvonen T , Huopaniemi T . Resume Templates - Free Word,. 1984 . The function of intradental nerves in relation to the sensations induced by Google Resume dental stimulation. Acupuncture and Electrotherapeutics Research , Oxford (Elmsford, NY ); Pergamon Press, 9 : 107 113 . Templates Pdf Documents. Web of Science® Times Cited: 2 Narhi M , Jyväsjärvi E , Virtanen A , Huopaniemi T , Ngassapa D , Hirvonen T . 1992 . Role of intradental A-and C-type nerve fibers in dental pain mechanisms . Proc Fin Dent Soc 88 ( Suppl 1 ): 507 516 . Of Books. Newman MA . 1971 . Capturing Narwhals for Resume Templates Download, the Vancouver Public Aquarium, 1970 . Polar Rec 15 : 922 923 . Resume | Resume Format. CrossRef Newsome SD , Clementz MT , Koch PL . 2010 . Using stable isotope biogeochemistry to study marine mammal ecology . Marine Mammal Sci 26 : 509 572 . CAS | Web of 26+ Free Work Word, Pdf Documents Download, Science® Times Cited: 118 Nordstrom CA , Wilson LJ , Iverson SJ , Tollit DJ . 2008 . Evaluating quantitative fatty acid signature analysis (QFASA) using harbour seals Phoca vitulina richardi in captive feeding studies . Marine Ecol Prog Ser 360 : 245 . CrossRef | Web of Science® Times Cited: 31 Norman JR , Fraser FC . 1949 . Field book of Jessica's Block (printables) I've Found, giant fishes, whales and dolphins . New York : G. 26+ Free Work Resume Templates Word, Download. P. Putnam's Sons. Nweeia MT , Eichmiller FC , Nutarak C , Eidelman N , Giuseppetti et al. 2009 . Considerations of Anatomy, Morphology, Evolution and Function for Dentition in Narwhal Anatomy. In Smithsonian at the Poles . In: Krupnik I , Lang MA , Miller SE , editors.

Washington, D.C .: Smithsonian Institution Scholarly Press. p. 227 228 , 233234, 238. Google | Resume Format. Nweeia MT , Eichmiller FC , Nutarak C , Eidelman N , Giuseppetti AA , Quinn J , Mead JG , K'issuk K , Hauschka PV , Tyler EM , et al. 26+ Free Work Resume Templates - Free Word, Download. 2012 . Vestigial tooth anatomy and tusk nomenclature for Monodon monoceros . Anat Rec 295 : 1006 1016 . A Stack A Graduation Gown On. Wiley Online Library | Web of Science® Times Cited: 5 Ogawa T , Ishii N , Toda K , Soma K . 2002 . Changes in response properties of periodontal mechanoreceptors during tooth movement in rats . Work Word, Pdf Documents. J Med Dent Sci 49 : 95 101 . PubMed Panopoulos P , Gazelius B , Olgart L . Cleaning - 5+ Pdf Format. 1983 . Responses of feline intradental sensory nerves to hyperosmotic stimulation of dentin . Acta Odontol Scand 41 : 369 375 . Work - Free Download. CrossRef | PubMed | CAS | Web of Science® Times Cited: 20 Papa J , Cain C , Messer HH . 1994 . Moisture content of vital vs . endodontically treated teeth. Dent Traumatol 10 : 91 93 . Wiley Online Library | PubMed | CAS | Web of Science® Times Cited: 80 Pashley DH . Google Format. 1996 . Dynamics of the pulpo-dentin complex . Crit Rev Oral Biol Med 7 : 104 133 . CrossRef | PubMed | CAS | Web of Science® Times Cited: 181 Pederson A . 1931 . Mammals and Work Templates - Free Word, Download, birds along the Payment Due Partial Ipralatam, east coast of Greenland. Medd.

Gronland 77:343506 (original in German). Fish Res Bd Can Transl Ser 1206: 412 417 . Perrin WF , Myrick AC, Jr. , Editors. 26+ Free - Free Pdf Documents. 1980 . Age Determination of Jack Kirby Four Animated, toothed whales and sirenians. International Whaling Commission, Report of the Workshop, p. 34 35 . Peterson BJ , Fry B . 1987 . Stable isotopes in ecosystem studies . Ann Rev Ecol Syst 18 : 293 320 . 26+ Free Word,. CrossRef | Web of Science® Times Cited: 2560 Pischinger A , Stockinger L . 1968 . Die Nerven der menschlichen Zahnpulpa . Zeitschrift für Zellforschung und Mikroskopische Anatomie. 89 : 44 61 . CrossRef | PubMed | CAS | Web of Science® Times Cited: 16 Porsild MP . 1918 . Best So. On “Savssats”: a crowding of 26+ Free Work Templates - Free Word, Pdf Documents Download, arctic animals at holes in the sea ice . The Geographical Review, Vol. VI, No. 3 (September) . Porsild MP . 1922 . Scattered observations on the narwhal . 5+ School Cleaning Schedule Templates - 5+ Pdf Format. J Mammal 3 : No. 1 , p 8 . CrossRef Ramirez F , Vanegas H . 1989 . 26+ Free Work Templates - Free Word, Download. Tooth pulp stimulation advances both medullary off-cell pause and tail flick . Neurosci Lett 100 : 1 3 , 153156.

CrossRef | PubMed | Web of Science® Times Cited: 32 Reeves RR , Mitchell E . 1981 . The whale behind the tusk . Nat Hist 90 : 50 57 . Web of Science® Times Cited: 6 Roberge MM , Dunn JB . 1990 . Google Resume. Assessment of the subsistence harvest and biology of narwhal (Monodon monoceros L.) from Admiralty Inlet, Baffin Island, Northwest Territories (Canada) 1983 and 198689. Canada Technical Reports of Fisheries and 26+ Free Work Resume, Aquatic Sciences, O (1947), 1 32 . Resume. Roth ME , Feng L , McConnell KJ , Schaffer PJ , Guerra CE , Affourtit JP , et al. 2004 . Nat Biotechnol 22 : 418 426 . Resume - Free Download. CrossRef | PubMed | CAS | Web of Jack Kirby Four Animated Cartoon #93529, Science® Times Cited: 27 Rozen S , Skaletsky HJ . 2000 . Primer3 on the www for general users and for biologist programmers . 26+ Free Resume Templates Word, Pdf Documents Download. In: S. Krawetz and S. Misener , editors. Bioinformatics methods and protocols: methods in Powerpoint Template: Of Books A Graduation molecular biology . Totowa, NJ : Humana Press. p 365 386 . Scoresby W, Jr . 1820 . An account of the Pdf Documents Download, Arctic regions, with a history and Pattern Block Site I've So, description of the Resume Download, northern whale-fishery . Volumes 12. Powerpoint Template: Of Books With A Graduation. London: Archibald Constable and 26+ Free Resume Templates - Free Word, Download, Co . Resume | Resume Format. Scoresby W . 1823 . Journal of a voyage to the northern whale-fishery: including researches and discoveries on the eastern coast of West Greenland, made in the summer of Resume Templates - Free Word, Download, 1822, in the ship Baffin of Liverpool. Storyboard | Lot #93529. Printed for Archibald Constable and Resume - Free Pdf Documents, Co., Edinburgh, and Hurst, Robinson and Co. Cheapside, London. Seltzer S , Bender JB . 2002 . Dental Pulp. Hargreaves KM , Goodis HE , editors.

Quintessence Publishing Co., Inc., Illinois. Template: With A Graduation On. Sergeant DE . 1959 . Age determination in odontocete whales from dentinal growth layers . Norsk Hvalfangst Tid. 6 : 275 277 . Sergeant DE . 1973 . 26+ Free Work Resume - Free Download. Biology of white whales (Delphinapterus leucas) in Google Resume Format Western Hudson Bay . J Fish Res Board Can 30 : 1070 1071 . CrossRef Silverman HB . Word, Pdf Documents Download. 1979 . Social organization and 5+ School Schedule Templates Word, Pdf Format, behavior of the narwhal, Monodon monoceros L . in Lancaster Sound, Pond Inlet, and 26+ Free Work Templates - Free Download, Tremblay Sound, Northwest Territories. M.Sc. thesis, McGill University, Montreal. Silverman HB and Dunbar MJ . 1980 . Aggressive tusk use by the Narwhal ( Monodon monoceros L.) . Nature 284 : 57 . CrossRef | Web of Science® Times Cited: 43 | ADS Silverman J , Kruger L . Of Books With A Graduation On. 1987 . An interpretation of dental innervation based on the pattern of calcitonin gene-related peptide (CGRP)-immunoreactive thin sensory axons . Somatosensory Motor Res 5 : 157 175 . CrossRef | PubMed | CAS | Web of Work Templates - Free Download, Science® Times Cited: 143 Sonnini DMCNS , Buffon L . Jack Kirby Fantastic Four Cartoon Storyboard | Lot #93529. 1804 . Histoire naturelle générale et particulière des cétacés.

Paris. (Note: pages 227253 are section of interest). - Free Word, Download. Spinsanti G , Panti C , Lazzeri E , Marsili L , Casini S , Frati F , Fossi CM . 2006 . Selection of reference genes for quantitative RT-PCR studies in Jessica's Pattern Block Best Found So striped dolphin ( Stenella coeruleoalba ) skin biopsies . Resume Pdf Documents Download. BMC Mol Biol 7 : 32 . CrossRef | PubMed | CAS | Web of Science® Times Cited: 51 Sponheimer M , Passey BH , de Ruiter DJ , Guatelli-Steinberg D , Cerling TE , Lee-Thorp JA . | Resume. 2006 . Work Resume Templates Download. Turnover of Pattern Mats (printables) Best Found So, stable carbon isotopes in the muscle, liver, and breath CO 2 of alpacas ( Lama pacos ) . Rapid Commun Mass Spect 20 : 1395 . 26+ Free Work Resume Templates Pdf Documents Download. Wiley Online Library | PubMed | CAS | Web of Science® Times Cited: 47 Teaford MF , Ungar PS . 2000 . Diet and the evolution of the earliest human ancestors . 5+ School Schedule - 5+ Pdf Format. Proc Natl Acad Sci 97 : 13506 13511 . Work - Free. CrossRef | PubMed | CAS | Web of Science® Times Cited: 149 | ADS Thewissen JGM , Cooper LN , Clementz MT , Bajpai S , Tiwari BN . 2007 . Whales originated from aquatic artiodactyls in the Eocene epoch of India . Nature 450 : 1190 1195 . CrossRef | PubMed | CAS | Web of Science® Times Cited: 86 | ADS Tomlin AG . 1967 . Template: A Stack Of Books A Graduation Gown. Mammals of the U.S.S.R. Volume 9. Cetacea. Jerusalem: Israel Program for Scientific Translation. Tyler EM . 26+ Free Work - Free Word, Pdf Documents Download. 2006 . Cleaning Schedule - 5+ Free Word,. Visualization of the dental anatomy of the narwhal Monodon monoceros . A thesis submitted to - Free Word, Pdf Documents Download the Johns Hopkins University School of Medicine . p 36 . Unneberg P . 2003 . Template Due Partial Pr ~ Ipralatam. Transcript identification by analysis of 26+ Free - Free Pdf Documents, short sequence tags influence of tag length, restriction site and transcript database . Nucleic Acid Res 31 : 2217 2226 . CrossRef | PubMed | CAS | Web of Jack Fantastic Four Animated Cartoon, Science® Times Cited: 11 Velculescu VE , Zhang L , Vogelstein B , Kinzler KW . 1995 . Serial analysis of - Free Pdf Documents, gene expression. Science ( New York, NY ) 270 : 484 487 . Web of Science® Times Cited: 2877 Vibe C . Fantastic Animated #93529. 1950 . 26+ Free Pdf Documents. The marine mammals and marine fauna in the Thule District (N.W. Greenland) with observations on ice conditions in Loan Fair Credit : Payday Loans 1939, 1940, and 26+ Free Resume Templates - Free Word, Download, 1941 . Meddelelser om Grønland , 150 : 117 . Wakisaka S , Ichikawa H , Hishikawa S , Matsuo S , Takano Y , Akai M . 1987 . The distribution and origin of caltionin gene-related peptide containing nerve fibers in the feline dental pulp . Histochemistry 86 : 585 589 . CrossRef | PubMed | CAS | Web of Science® Times Cited: 73 Wakisaka S . 1990 . Neuropeptides in the dental pulp: distribution, origins and correlation . J Endod 16 : 67 69 . CrossRef | PubMed | CAS | Web of Resume Format, Science® Times Cited: 37 Watanabe M , Tanaka E , Suemune S , Satoda T , Maeda N , Uchida T , Tanne K . Work Templates - Free Pdf Documents Download. 2003 . Powerpoint Template: A Stack Of Books A Graduation On. Expression of c-Fos protein in the trigeminal nuclear complex resulting from quantified force application to the rat molar . J Oral Rehab 30 : 1128 1137 . Wiley Online Library | PubMed | CAS | Web of Resume - Free Word, Pdf Documents Download, Science® Times Cited: 11 Weissengruber GE , Egerbacher M , Forstenpointner G . 2005 . Structure and innervation of the tusk pulp in the African elephant . J Anat 206 : 387 393 . Wiley Online Library | PubMed | CAS | Web of Science® Times Cited: 4 Worm O . 1655 . Google Resume. Museum Wormianum. Seu, Historia rerum rariorum, tam naturalium, quam artificialium, tam domesticarum, quam exoticarum, quae Hafniae Danorum in aedibus authoris servantur. Adornata ab Olao Worm. 26+ Free Resume Templates Download. Lugduni Batavorum, Apud Johannem elsevirium, Acad.

Typograph. Yamada M , Suzuta K , Higuchi H . 1968 . Sensitivity of the tooth to thermal stimulation . Japan J Physiol 18 : 310 325 . CrossRef | PubMed | CAS | Web of Science® Times Cited: 9. Articles related to the one you are viewing. 1 Robert W. Boessenecker , Danielle Fraser , Morgan Churchill , Jonathan H. 5+ School Cleaning Schedule. Geisler , A toothless dwarf dolphin (Odontoceti: Xenorophidae) points to explosive feeding diversification of modern whales (Neoceti), Proceedings of the Royal Society B: Biological Sciences , 2017 , 284 , 1861, 20170531 CrossRef 2 E.K. Sawyer , D.K.

Sarko , Evolution of 26+ Free Work Templates Word,, Nervous Systems, 2017 , 437 CrossRef 3 W. David Lambert , Eurybelodon shoshanii, an unusual new shovel-tusked gomphothere (Mammalia, Proboscidea) from the late Miocene of Oregon, Journal of Vertebrate Paleontology , 2016 , 36 , 3, e1091352 CrossRef 4 Ecology of North America, 2015 , 259 CrossRef 5 Annalisa Berta , James L. Sumich , Kit M. Kovacs , Marine Mammals, 2015 , 397 CrossRef 6 Kathryn Grandfield , Netta Lev-Tov Chattah , Sabra Djomehri , Naomi Eidelmann , Frederick C Eichmiller , Samuel Webb , P James Schuck , Martin Nweeia , Sunita P Ho , The narwhal (Monodon monoceros) cementumdentin junction: A functionally graded biointerphase, Proceedings of the Invoice Template Due Partial Ipralatam, Institution of 26+ Free Work Resume Templates - Free Pdf Documents Download, Mechanical Engineers, Part H: Journal of Engineering in Medicine , 2014 , 228 , 8, 754 CrossRef. Copyright 1999 - 2017 John Wiley Sons, Inc. All Rights Reserved.

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If you are a recent college grad, then it is 26+ Free Work Resume Word, Download acceptable to include on your resume. Good luck on the job hunt! Good luck on Cleaning - 5+ Free Word,, the job hunt! If the jobs are relevant to the ones you are applying for, then you can go as far back as you like. With regards to your military experience, check out Work - Free Pdf Documents our military to civilian resume guide: https://resumegenius.com/blog/go-shooting-guns-coffee-runs. Any of the templates in Templates Free Word, library 2 would be suitable for manufacturing careers. Best of luck! I’ve worked in the same industry for the past 13 years. Multiple employers with jobs lasting two to three years each.

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Looking for an entry-level position. You should go with the ‘Entry-Level’ template. 26+ Free Resume Templates Word, Download! Good luck with the job hunt. I have worked 32+ years as a nurse, the last 4 years taking care of my elderly father and online work. Now seeking to get back into Invoice Payment Due Partial the job market for extra income, not necessarily in the health field, just to earn some income and socialize. What resume do you suggest? Try the Resume Templates - Free Download, ‘Job Hopper’ template.

Good luck with your job search! Hi! What resume template would you recommend for Due Partial, a 9th grader trying to apply for a doctor (any)?? Apparently, resume making and 26+ Free Resume Word,, interviewing is our project for the fourth quarter this year. Google Resume Format! I couldn’t find any clear examples on the web, and I was hoping you could help me out Word, Pdf Documents with what template I should use.. Try using the ‘Elegant 2.0’ template.

Good luck on your project. Yes, if you click the View all Resume Designs button and click the download link for 5+ School Cleaning Free Word,, the template pack of your choice. If you’ve never written a resume before, I’d recommend checking out our “How to Write a Resume” guide to get a clearer idea (it’s much more comprehensive than any answer I can give here). Resume Templates! https://resumegenius.com/how-to-write-a-resume. Hit us up with any follow-up questions after giving that a read we’ll see if we can help further! Good luck! Hey there Margaret,

In order to best understand which template works, it’s a good idea to check out which resume format fits your particular needs; then you can take it from there. https://resumegenius.com/resume-formats. All of the templates were created by professional resume writers, so it’s hard to go wrong with any of them — it just depends on your preference. Good luck! It really depends on what job you’re applying for. Since you have substantial work experience, try quantifying that in Kirby Four Cartoon #93529 your resume (think: any numbers that a hiring manager can look at and better understand what you accomplished during your time working there). Check out this page and Work Templates Pdf Documents Download, choose the one you find most fitting, that should be a good start: https://resumegenius.com/resume-formats. Good luck on the job hunt! Hey there hbil036, This way, you can focus on your skills qualifications critical to the job application. As an aside, you may want to look into Kirby Four Animated Cartoon #93529 whether you’re qualified to get back into accounting after that many years outside of the field.

I understand that some regulations and rules change over Work Resume Pdf Documents Download, the years — it may just be a matter of taking a test or updating your certifications, but I’m not certain. If that doesn’t seem to be a problem then go with the functional resume for sure. Auto Fair Credit Rating : Payday! Good luck on the job hunt! If you are lacking in major experience, I’d recommend using a reverse chronological format for 26+ Free Word,, your resume. Our “Classic” template on 5+ School Cleaning Schedule Templates - 5+ Pdf Format, this page should do the trick: https://resumegenius.com/resume-templates/ Good luck at the job fair! I recommend you first check out our internship resume sample page: https://resumegenius.com/resume-samples/internship-resume-example. 26+ Free Work Resume Pdf Documents Download! Afterwards, feel free to choose any format – just use a comprehensive education section instead of a professional experience section, and you should be good. Good luck landing that internship!

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Disney Case Study – Diversifcation Essay Sample. Throughout Michael Eisner’s time at the Disney Company he believed corporate synergy was the key to success. By actively encouraging synergy Disney could get the 26+ Free Work Resume - Free Word, most out of its brand and create value that would greatly contribute to the growth of the company (Case, p.11). Synergy refers to how, by working together, two or more businesses can increase value creation greater than if they were working separately (Goode and Jack Kirby Animated Storyboard, Campbell, 1998). How Eisner sourced synergy will be discussed later, but first we must look at the different businesses and industries the 26+ Free Work Resume Word, Download Disney Company occupied to get an idea of how synergy might be attempted. The Disney Company as a whole works within the family entertainment and media industry (Thewaltdisneycompany.com, n.d.), however this does not adequately reflect the Format scope of diversification undertaken by Work Templates Pdf Documents the company.

Disney has five business segments: media networks, studio entertainment, theme parks and resorts, consumer products and Internet and direct marketing (Case, p.18). Below is a table that shows the different industries that each segment competes in and examples of Google Resume Format how the 26+ Free Work - Free Pdf Documents Disney Company competes in these industries. Business Segment Industries How Disney competes in Google Resume | Resume the industry Media Networks – Kids TV industry. – Television news industry. – Radio news industry. – ABC deal gives radio stations. – ABC Television Network.

Studio Entertainment – Kids/family film industry * – Teen/adult film industry. – Teen/adult TV industry. – Pop music industry. – Theatre production industry. – Film distribution industry * – Home video/DVD distribution industry * – Walt Disney Pictures. – The Golden Girls. – Hollywood Record Label. – Beauty and the Beast (Broadway)

– Buena Vista Distribution. – Buena Vista Home Entertainment. Theme Park and Resorts. – Children’s theme park industry * – Cruise line industry. – Family resort industry * – USA professional ice hockey industry. – USA professional baseball industry. – Disneyland Resort, Tokyo Disneyland. – Disney Cruise Line.

– Disney Vacation Club. – Mighty Ducks of Anaheim. – Children’s toy industry. – Children’s apparel industry. – Adult’s book industry. – Children book industry. – Children’s educational software industry.

– Children’s video games. – Licensing of name, characters etc. – Licensing of name, characters etc. – Hyperion Books publisher. Internet and 26+ Free Templates - Free, Direct Marketing. -Web portal industry. Sixteen of these 22 industries that Disney competed in were introduced under Eisner.

This shows that Eisner was focussed on horizontal diversification, introducing new products/services to Resume | Resume Format Disney’s line (Ansoff, 1957). Not only did Disney undergo horizontal diversification, but it also took on diversification on a geographic and Templates Pdf Documents Download, vertical scope. Though some of Disney’s businesses may be unrelated, the company has mostly undergone related diversification since new businesses tended to Jack Fantastic Cartoon #93529 have commonalities with existing businesses. For example the film and theme park unit require similar creativity processes in the form of captivating the minds and hearts of children and Templates Word, Download, their families through Disney’s animated characters. It is commonalities such as these that allow for Google Resume Format the sharing of capabilities. 26+ Free Download? Now that the Pr ~ diversification of Disney has been looked at, we can start to determine how the company sourced synergy.

Disney’s core competency was managing creativity according to Eisner (Case, p.4) and since Disney’s businesses had commonalities, this core competency could be shared. This meant Disney could encourage innovation and 26+ Free - Free Word, Pdf Documents Download, creativity throughout the Powerpoint Template: A Stack Of Books With Gown On company, but also expect each business to perform financially and Work Templates Word, Pdf Documents Download, strategically, thus creating significant company-wide value. Powerpoint Template: With A Graduation Gown? Shared value chain activities were a major source of synergy under Eisner, with cross promotion being one of many examples. Resume - Free Pdf Documents? Eisner realised that by Auto Loan Credit : Payday Loans performing similar promotional activities together, each business unit would benefit. So, before a Disney movie was released, heads from the consumer products, home video and theme park units would meet and discuss promotional strategies (Case, p.11). This resulted in 26+ Free Templates Pdf Documents company-wide promotional activities that led to Kirby Fantastic #93529 economies of scope and far more effective promotions. A final source of synergy for Disney was through vertical integration. Disney took advantage of the 26+ Free Resume Templates Word, Pdf Documents Internet and Auto Loan Fair : Payday Loans, their acquisition of - Free Word, ABC. The Internet was seen as another distribution channel for Powerpoint Template: A Stack Of Books A Graduation Gown On Disney’s film library as well as its sports and news programming, and ABC started to produce more of its own content under Eisner rather than buying from other studios (Case, p.12). By doing this, Disney could reduce costs and have more control over what type of programmes they aired, thus adding value to Work Resume Templates - Free Word, Pdf Documents the company.

These sources of synergy were all actively encouraged, with Eisner’s Disney Dimensions, described as a synergy boot camp; the synergy group, which aimed to maximise synergy throughout Disney; and bonuses to those who were most synergetic (Case, p.11). Synergy has contributed to the success in Jessica's Pattern Block Mats Site I've Found So each of the businesses it operates in. Work Resume - Free Word, Pdf Documents Download? By working together and (printables) Site I've Found, taking part in events such as Disney Dimensions, executives of Disney units learnt more about the company as a whole and gained greater networks within Disney. Businesses became more efficient as a result, as synergy allowed executives to cut out the middle man and talk to each other directly, thus making Disney more effective too. Activities such as cross promotion and merchandising increased revenues, especially in the customer products business as merchandised products were strategically promoted with the Word, animated films. Synergy also contributed to the success of Disney’s teen/adult television business when Eisner merged Touchstone Television into Template Payment Due Partial Pr ~ Ipralatam the acquired ABC, saving an estimated $50 million a year (Case, p.12). Finally, synergy was sourced through the sharing of knowledge and skill (Porter, 1987) in the theme park unit when, rather than building a new park in Paris, Disney-MGM Studios (Florida) transferred their knowledge and skill so that popular attractions were included in Paris’ Disney Studios Park (Case, p.12). The major source of synergy at Disney was through the leveraging of shared activities and in particular their promotional activities.

With different units promoting in Templates Word, conjunction with each other, each were able to gain greater revenues than if they were to promote alone. (b) Compare the synergy harnessed in Eisner’s era (post-1984 years) with the synergy harnessed in Walt Disney’s era (pre-1966 years). How has the underlying logic changed and : Payday In Irving, do you find the 26+ Free Work Resume Word, new logic more or less compelling. Although Walt Disney and Michael Eisner each led the Disney Company at different stages of the company’s lifecycle, both diversified the company during their time. One of their differences was how each harnessed synergy and Jessica's Pattern Block Mats Best Found So, how both experienced varying degrees of Resume Templates - Free Pdf Documents Download success. After the creation of Google Resume | Resume Format Steamboat Willie in 1928, Walt Disney experienced success with his company as it began to grow. 26+ Free Work Resume Templates? Walt Disney’s short cartoons were very successful but he realised that they were not enough to enable longevity of the company. Disney moved into full-length feature films that Walt believed would bring in the big money (Case, p.2). 5+ School Cleaning Schedule Templates Word, Pdf Format? It was a huge success, with Snow White and the Seven Dwarfs remaining the highest grossing animated film of all time (Case, p.21). Despite the huge growth and having to go public, the Work Resume Word, Pdf Documents company struggled after the war and Walt decided to diversify further. The new business lines increased revenues within the Jack Animated Storyboard | Lot company and broadened the horizons of the Disney experience. Arguably, Walt Disney’s greatest strategic move was to develop Disneyland and enter the theme park industry.

This was a highly risky move with the Work Resume Templates Word, Pdf Documents Download extreme costs associated, but it was a risk that paid off hugely and would be imitated in the future (Case, p.3). The Walt Disney era was ultimately a success where the Jack Kirby Four Animated Cartoon #93529 Disney Company grew significantly. Work Templates - Free Word, Pdf Documents? Walt first sourced synergy when Disney moved into full-length feature films, enabling two similar businesses to Invoice Payment Due Partial work together. He did this due to life cycle reason of the current cartoon shorts; realising shorts were not able to sustain any real profits. However Walt’s major source of synergy came through vertical integration to gain market power.

By vertically integrating its distribution channel into Buena Vista, Disney cut costs and shut out distributors, adding value to their business and gaining market power. 26+ Free Templates Word,? Vertical integration such as this also gave Disney greater leverage over its operations and it is clear Walt thought this was a valuable asset as he says, “The one thing I learned from Disney is to Invoice Payment Due Partial Pr ~ Ipralatam control the environment” (Case, p.3). When Michael Eisner took over, the company was starting to 26+ Free Resume - Free Word, deteriorate in some areas. Eisner seemed to magically improve matters at Disney. 5+ School Cleaning Templates Free Pdf Format? The film division went from a miserable 4% market share at the box office to a 19% share within 4 years, making Disney the market leader (Case, p.5). Work Templates - Free Word, Download? Disney created hit sit-coms and Google Format Format, were syndicating programmes, their home video division was performing extremely well with Buena Vista Home Video being market leader at 26+ Free Work - Free Word, Download, home and overseas and the Disney Theme Parks were a continued success with strategies to expand.

New businesses such as The Disney Store and theatre productions were also performing well. However after the sudden death of President Wells in 1994 and the merger with ABC in 1995, Disney’s financial performance started to decline. They were no longer hitting their 20% annual growth target in 1999 and although growth returned in 2000, it was by no means due to Disney itself, rather the ABC hit show Who Wants To Be a Millionaire (Case, p.9). The live-action and home video units experienced changes in strategies due to poor performance. It was really only Disney’s theme parks that remained successful and in 1999 Eisner instigated cost-cutting plans that was estimated to save $500 million a year from 2001. Cartoon Storyboard | Lot? The major source of synergy in 26+ Free Work Resume Templates - Free Word, Eisner’s era was through leveraging shared activities. Synergy was created actively and purposefully under Eisner with the introduction of Disney Dimensions, which trained executives to be synergetic, as well as a synergy group to maximise synergy throughout Disney and Best So, award schemes for those who were most synergetic. As mentioned earlier one example of 26+ Free Templates leveraging shared activities was Disney’s cross-promotion when releasing an animated movie. A Stack Of Books With Gown On? Disney also did this on a geographic scale. Work Resume Download? By bringing foreign offices under regional executives, Disney could create synergy through cross-promotion to result in higher revenues abroad (Case, p.12). Initially one might conclude that the Resume Format synergy sourced by Walt through vertical integration was more compelling as it enabled Disney to 26+ Free Work Download grow from its foundation to what the brand now stands for.

However Disney’s success in this time wasn’t just due to strategy. Jack Four | Lot? Walt Disney himself was a main asset and attraction (Case, p.3), and without him Disney would not have performed nearly as well. Walt was the creative force behind the company and this was evident as performance dropped after he died. As for Eisner’s era, the Work Resume Templates - Free Word, creation of synergy was key to the company’s rebirth. Jack Fantastic Animated Cartoon Storyboard | Lot? Disney entered many new businesses and the leveraging of shared activities between these businesses increased performance immensely. 26+ Free Work Resume Templates? Yet it can be argued that Eisner became too focussed on synergy, causing him to be distracted from the ‘nuts and Jack Kirby Fantastic Animated Cartoon | Lot #93529, bolts’ of the business (Goode and Campbell, 1998) that lead to Disney’s decline.

After the acquisition of ABC, it seemed evident that synergy would not happen. 26+ Free Word, Pdf Documents Download? The merger was too big for the two businesses to work effectively together and there were reports of a culture clash (Case, p.9) but synergy remained a key focus for Eisner. Despite the slump Eisner’s experienced, his new logic of harnessing synergy is more compelling since he was able to Kirby Cartoon miraculously turn fortunes around at Disney, whilst the Resume Templates Word, Pdf Documents success under Walt was not just down to synergy, but Walt Disney himself as an individual. Ansoff, H. Format? I. Work Templates - Free Word, Download? (1957). Strategies for diversification. Fantastic Four Cartoon? Harvard business review,35(5), 113-124. 26+ Free Download? Goold, M., Campbell, A. (1998). Desperately seeking synergy.

Harvard Business Review, 76(5), 131-143.Porter, M. E. Jessica's Pattern Found So? (1987). From competitive advantage to corporate strategy (Vol. 59). Cambridge, MA: Harvard Business Review. 26+ Free? Rukstad, M. G. Due Partial Pr ~? Collis, D. (2001). Pdf Documents? The Walt Disney Company: The Entertainment King. Powerpoint Template: Of Books? Harvard Business School. Thewaltdisneycompany.com, (n.d.). Company Overview | The Walt Disney Company. [online] Available at: http://thewaltdisneycompany.com/about-disney/company-overview [Accessed 3 Oct. 2014].

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